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Enhancers activate from afar Nat. Cell Biol. (IF 17.3) Pub Date : 2025-02-13 Sabrya Carim
Enhancers control the expression of their target genes from a distance, and many genes are controlled by multiple enhancers. However, how the individual enhancer elements interact and cooperate is poorly understood. A study by Thomas, Feng et al. designed a reporter system that enables enhancer–enhancer cooperativity across distances in the mammalian genome to be studied. The authors found that the
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Mesoderm FN1 shaping the heart Nat. Cell Biol. (IF 17.3) Pub Date : 2025-02-13 Daryl J. V. David
Cardiac development involves tissue mechanics, cell polarity, growth and cell adhesion but their precise co-ordination for proper morphogenesis in mammals is unclear. Arriagada et al. report that the extracellular matrix (ECM) protein fibronectin (FN1) synthesized by the mesoderm affects multiple cell behaviours in the dorsal pericardial wall of the developing mouse heart. The authors used Fn1flox/−;Mesp1Cre
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Restoring placental methylome Nat. Cell Biol. (IF 17.3) Pub Date : 2025-02-13 Stylianos Lefkopoulos
Human trophoblast stem cells (hTSCs) are used to model placentation, but whether their epigenome recapitulates primary trophoblast remains underexplored. A new study shows that hTSCs display an atypical methylome compared with trophectoderm and first trimester cytotrophoblast (CTB), which can be restored by DNMT3L overexpression. Lea et al. performed whole-genome bisulphite sequencing on cultured hTSCs
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Ribophagy relies on Rpl12 Nat. Cell Biol. (IF 17.3) Pub Date : 2025-02-11 Katarzyna Tutak, Katrin Karbstein
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Rpl12 is a conserved ribophagy receptor Nat. Cell Biol. (IF 17.3) Pub Date : 2025-02-11 Yuting Chen, Jiaxin Hu, Pengwei Zhao, Jie Fang, Yingqi Kuang, Zhaojie Liu, Shuling Dong, Weijing Yao, Yuanyuan Ding, Xinhui Wang, Yibin Pan, Jianbin Wu, Jingwei Zhao, Jing Yang, Zhenzhong Xu, Xiaodi Liu, Yi Zhang, Choufei Wu, Liqin Zhang, Mingzhu Fan, Shan Feng, Zhi Hong, Zhangming Yan, Hongguang Xia, Kaiyue Tang, Bing Yang, Wei Liu, Qiming Sun, Kunrong Mei, Wei Zou, Yunpeng Huang, Du Feng, Cong Yi
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Regulating MYC translation in cancer Nat. Cell Biol. (IF 17.3) Pub Date : 2025-02-07 Zainab Hussain, Mara H. Sherman
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A quantitative ultrastructural timeline of nuclear autophagy reveals a role for dynamin-like protein 1 at the nuclear envelope Nat. Cell Biol. (IF 17.3) Pub Date : 2025-02-07 Philip J. Mannino, Andrew Perun, Ivan V. Surovtsev, Nicholas R. Ader, Lin Shao, Elisa C. Rodriguez, Thomas J. Melia, Megan C. King, C. Patrick Lusk
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Time for lipid cell biology Nat. Cell Biol. (IF 17.3) Pub Date : 2025-02-05 Xiao-Wei Chen, Anthony S. Don, Maria Fedorova, Takeshi Harayama, Natalie Krahmer, Shigekazu Nagata, Priyanka Narayan, Dequina Nicholas, Sara M. Nowinski, Yasunori Saheki, Clay F. Semenkovich, Xiaoai Zhao, Yilong Zou
It is an exciting time for lipid metabolism and membrane cell biologists as technological progress has increased our ability to study lipids in cells. We asked leaders studying lipid cell biology from different perspectives to share what questions they are most interested in and what tools they believe the field is currently lacking.
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Functional screen identifies RBM42 as a mediator of oncogenic mRNA translation specificity Nat. Cell Biol. (IF 17.3) Pub Date : 2025-02-04 Joanna R. Kovalski, Goksu Sarioglu, Vishvak Subramanyam, Grace Hernandez, Gilles Rademaker, Juan A. Oses-Prieto, Macey Slota, Nimmy Mohan, Kaylee Yiakis, Isabelle Liu, Kwun Wah Wen, Grace E. Kim, Sohit Miglani, Alma L. Burlingame, Hani Goodarzi, Rushika M. Perera, Davide Ruggero
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How a ribosomal protein avoids mixed signals Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-31 Kanda M. Borgognoni, Robin E. Stanley
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An avoidance segment resolves a lethal nuclear–mitochondrial targeting conflict during ribosome assembly Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-31 Michaela Oborská-Oplová, Alexander Gregor Geiger, Erich Michel, Purnima Klingauf-Nerurkar, Sven Dennerlein, Yury S. Bykov, Simona Amodeo, André Schneider, Maya Schuldiner, Peter Rehling, Vikram Govind Panse
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SNORD113–114 cluster maintains haematopoietic stem cell self-renewal via orchestrating the translation machinery Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-31 Hui Wang, Zhaoru Zhang, Chenxi Han, Penglei Jiang, Jiayue Xu, Yingli Han, Deyu Huang, Jian Li, Jie Zhou, Michael Durnin, Shiyuan Chen, Yaxin Liu, Jinghao Sheng, Jie Cao, Jianzhao Liu, Bing Liu, Jia Yu, Fang Wang, Pengxu Qian
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Cytoplasmic flow is a cell size sensor that scales anaphase Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-31 Olga Afonso, Ludovic Dumoulin, Karsten Kruse, Marcos Gonzalez-Gaitan
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Multiomics reveals that triacylglycerol mobilization helps drive recovery from mitochondrial stress Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-24
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Mitochondrial protein import stress Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-22 Nikolaus Pfanner, Fabian den Brave, Thomas Becker
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Tripotency in human pancreas Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-16 Stylianos Lefkopoulos
Research in the mouse indicates that all three pancreas epithelial compartments derive from a single and transient progenitor. Now, a study of the human fetal pancreas identifies a leucine-rich repeat-containing G-protein-coupled receptor 5-positive (LGR5+) cell population capable of generating all exocrine and endocrine pancreatic lineages. Andersson-Rolf et al. derived pancreas organoid lines from
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Chaperoning RNA into granules Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-16 Daryl J. V. David
Biomolecular condensate formation requires multivalent interactions and rapid turnover. Stress granules are condensates formed through RNA–protein interactions under stress, requiring the RNA-binding protein G3BP. Parker et al. show that G3BP1 is an RNA condenser and promotes initial RNA–RNA interactions but is dispensable for stability; instead, stability requires RNA–RNA interactions. The authors
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Real-time mirroring of therapy Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-16 Stylianos Lefkopoulos
Early assessment of treatment efficacy can be invaluable in cases of aggressive tumours, such as glioblastoma. Logun et al. generated glioblastoma organoids (GBOs) from patients participating in a chimeric antigen receptor (CAR) T cell clinical trial to monitor the therapeutic response in real time. The authors obtained tumour specimens from six patients with glioblastoma enrolled in a phase 1 CAR
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Battling Diamond-Blackfan anaemia Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-16 Stylianos Lefkopoulos
Gene therapy strategies against Diamond-Blackfan anaemia (DBA) have been hampered by the multiple and heterogeneous causative mutations. A study now shows that modulating GATA1 expression is sufficient to tackle the erythroid maturation arrest in DBA models and patient-derived samples. Voit et al. first identified endogenous regulatory elements (hG1E-GATA1) guiding erythroid-restricted expression of
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Adapting to improve the author experience Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-16
Here we discuss approaches to refine our editorial processes.
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Retraction Note: Tumour microenvironment programming by an RNA–RNA-binding protein complex creates a druggable vulnerability in IDH-wild-type glioblastoma Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-16 Lele Wu, Zheng Zhao, Yong Jae Shin, Yiyun Yin, Anandhkumar Raju, Thamil Selvan Vaiyapuri, Khaireen Idzham, Miseol Son, Yeri Lee, Jason K. Sa, Joelle Yi Heng Chua, Bilal Unal, You Zhai, Wenhua Fan, Lijie Huang, Huimin Hu, Jayantha Gunaratne, Do-Hyun Nam, Tao Jiang, Vinay Tergaonkar
Retraction to: Nature Cell Biology (2024) 26:1003-1018 https://doi.org/10.1038/s41556-024-01428-5
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Mitotic lethality prevents inflammation Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-14 Christian Zierhut, Andreas Villunger
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Homologous recombination promotes non-immunogenic mitotic cell death upon DNA damage Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-13 Radoslaw Szmyd, Sienna Casolin, Lucy French, Anna G. Manjón, Melanie Walter, Léa Cavalli, Christopher B. Nelson, Scott G. Page, Andrew Dhawan, Eric Hau, Hilda A. Pickett, Harriet E. Gee, Anthony J. Cesare
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Spatial transcriptomic characterization of a Carnegie stage 7 human embryo Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-10 Lina Cui, Sirui Lin, Xiaolong Yang, Xinwei Xie, Xiaoyan Wang, Nannan He, Jingyu Yang, Xin Zhang, Xiaojian Lu, Xiaodi Yan, Yifei Guo, Bailing Zhang, Ran Li, Hefan Miao, Mei Ji, Runzhao Zhang, Leqian Yu, Zhenyu Xiao, Yulei Wei, Jingtao Guo
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Boosting cargo turnover with receptor mobility Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-09 Yi Lu, Chunmei Chang
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MDM2 functions as a timer reporting the length of mitosis Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-09 Luke J. Fulcher, Tomoaki Sobajima, Caleb Batley, Ian Gibbs-Seymour, Francis A. Barr
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The nuclear matrix stabilizes primed-specific genes in human pluripotent stem cells Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-09 Gang Ma, Xiuling Fu, Lulu Zhou, Isaac A. Babarinde, Liyang Shi, Wenting Yang, Jiao Chen, Zhen Xiao, Yu Qiao, Lisha Ma, Yuhao Ou, Yuhao Li, Chen Chang, Boping Deng, Ran Zhang, Li Sun, Guoqing Tong, Dongwei Li, Yiming Li, Andrew P. Hutchins
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Transcription factor networks in cellular quiescence Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-09 Mithun Mitra, Sandra L. Batista, Hilary A. Coller
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Precursor occupancy controls mitochondrial import channel via proteolysis Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-08
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FAT1 alterations contribute to chromosomal instability in cancer cells Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-08
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H3K36 methylation regulates cell plasticity and regeneration in the intestinal epithelium Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-08 Alison R. S. Pashos, Anne R. Meyer, Cameron Bussey-Sutton, Erin S. O’Connor, Mariel Coradin, Marilyne Coulombe, Kent A. Riemondy, Sanjana Potlapelly, Brian D. Strahl, Gunnar C. Hansson, Peter J. Dempsey, Justin Brumbaugh
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Mapping the developmental trajectory of human astrocytes reveals divergence in glioblastoma Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-08 Caitlin Sojka, Hsiao-Lin V. Wang, Tarun N. Bhatia, Yangping Li, Pankaj Chopra, Anson Sing, Anna Voss, Alexia King, Feng Wang, Kevin Joseph, Vidhya M. Ravi, Jeffrey Olson, Kimberly Hoang, Edjah Nduom, Victor G. Corces, Bing Yao, Steven A. Sloan
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Triacylglycerol mobilization underpins mitochondrial stress recovery Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-08 Zakery N. Baker, Yunyun Zhu, Rachel M. Guerra, Andrew J. Smith, Aline Arra, Lia R. Serrano, Katherine A. Overmyer, Shankar Mukherji, Elizabeth A. Craig, Joshua J. Coon, David J. Pagliarini
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Chromosome mis-segregation triggers cell cycle arrest through a mechanosensitive nuclear envelope checkpoint Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-08 Solène Hervé, Andrea Scelfo, Gabriele Bersano Marchisio, Marine Grison, Kotryna Vaidžiulytė, Marie Dumont, Annapaola Angrisani, Adib Keikhosravi, Gianluca Pegoraro, Mathieu Deygas, Guilherme P. F. Nader, Anne-Sophie Macé, Matteo Gentili, Alice Williart, Nicolas Manel, Matthieu Piel, Yekaterina A. Miroshnikova, Daniele Fachinetti
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Gallbladder-derived retinoic acid signalling drives reconstruction of the damaged intrahepatic biliary ducts Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-08 Jianbo He, Shuang Li, Zhuolin Yang, Jianlong Ma, Chuanfang Qian, Zhuofu Huang, Linke Li, Yun Yang, Jingying Chen, Yunfan Sun, Tianyu Zhao, Lingfei Luo
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Mitochondrial YME1L1 governs unoccupied protein translocase channels Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-07 Meng-Chieh Hsu, Hiroki Kinefuchi, Linlin Lei, Reika Kikuchi, Koji Yamano, Richard J. Youle
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Phase separation of initiation hubs on cargo is a trigger switch for selective autophagy Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-07 Mariya Licheva, Jeremy Pflaum, Riccardo Babic, Hector Mancilla, Jana Elsässer, Emily Boyle, David M. Hollenstein, Jorge Jimenez-Niebla, Jonas Pleyer, Mio Heinrich, Franz-Georg Wieland, Joachim Brenneisen, Christopher Eickhorst, Johann Brenner, Shan Jiang, Markus Hartl, Sonja Welsch, Carola Hunte, Jens Timmer, Florian Wilfling, Claudine Kraft
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Metabolic rewiring in skin epidermis drives tolerance to oncogenic mutations Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-06 Anupama Hemalatha, Zongyu Li, David G. Gonzalez, Catherine Matte-Martone, Karen Tai, Elizabeth Lathrop, Daniel Gil, Smirthy Ganesan, Lauren E. Gonzalez, Melissa Skala, Rachel J. Perry, Valentina Greco
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Speckle signatures dictate cancer prognosis Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-03 Hiro Takakuwa, Tetsuro Hirose
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A nucleolar mechanism suppresses organismal proteostasis by modulating TGFβ/ERK signalling Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-03 Huadong Zhu, Reut Bruck-Haimson, Adam Zaretsky, Irit Cohen, Roni Falk, Hanna Achache, Yonatan B. Tzur, Ehud Cohen
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Chaperone-mediated insertion of mitochondrial import receptor TOM70 protects against diet-induced obesity Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-03 Pedro Latorre-Muro, Tevis Vitale, Matthew Ravichandran, Katherine Zhang, Jonathan M. Palozzi, Christopher F. Bennett, Arantza Lamas-Paz, Jee Hyung Sohn, Thomas D. Jackson, Mark Jedrychowski, Steven P. Gygi, Shingo Kajimura, Anna Schmoker, Hyesung Jeon, Michael J. Eck, Pere Puigserver
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AMPK-regulated glycerol excretion maintains metabolic crosstalk between reductive and energetic stress Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-02 Xuewei Zhai, Ronghui Yang, Qiaoyun Chu, Zihao Guo, Pengjiao Hou, Xuexue Li, Changsen Bai, Ziwen Lu, Luxin Qiao, Yanxia Fu, Jing Niu, Binghui Li
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Nuclear speckles regulate functional programs in cancer Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-02 Katherine A. Alexander, Ruofan Yu, Nicolas Skuli, Nathan J. Coffey, Son Nguyen, Christine L. Faunce, Hua Huang, Ian P. Dardani, Austin L. Good, Joan Lim, Catherine Y. Li, Nicholas Biddle, Eric F. Joyce, Arjun Raj, Daniel Lee, Brian Keith, M. Celeste Simon, Shelley L. Berger
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The transcription factor GABPA is a master regulator of naive pluripotency Nat. Cell Biol. (IF 17.3) Pub Date : 2025-01-02 Chengjie Zhou, Meng Wang, Chunxia Zhang, Yi Zhang
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TRACERx analysis identifies a role for FAT1 in regulating chromosomal instability and whole-genome doubling via Hippo signalling Nat. Cell Biol. (IF 17.3) Pub Date : 2024-12-30 Wei-Ting Lu, Lykourgos-Panagiotis Zalmas, Chris Bailey, James R. M. Black, Carlos Martinez-Ruiz, Oriol Pich, Francisco Gimeno-Valiente, Ieva Usaite, Alastair Magness, Kerstin Thol, Thomas A. Webber, Ming Jiang, Rebecca E. Saunders, Yun-Hsin Liu, Dhruva Biswas, Esther O. Ige, Birgit Aerne, Eva Grönroos, Subramanian Venkatesan, Georgia Stavrou, Takahiro Karasaki, Maise Al Bakir, Matthew Renshaw, Hang
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Bridging art and science Nat. Cell Biol. (IF 17.3) Pub Date : 2024-12-20 Sabrya Carim
Ahna Skop is a geneticist and artist at the University of Wisconsin–Madison, USA. Ahna’s lab studies the assembly and function of mammalian midbody and midbody remnant, which are assembled at the end of mitosis. Ahna also leans into her passion for scientific art to engage the public. We were fascinated to hear Ahna’s thoughts on open questions and challenges in the field and to learn about her passion
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Misshapen chromosomes in check by mechanics Nat. Cell Biol. (IF 17.3) Pub Date : 2024-12-18 Joana C. Macedo, Maria M. da Silva, Elsa Logarinho
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Cu in cancer metabolism Nat. Cell Biol. (IF 17.3) Pub Date : 2024-12-10 Zhe Wang
As an essential metal co-factor, Cu accumulation has been linked with tumorigenesis, but the underlying mechanism remains to be fully defined. A study now reports that Cu, as a subunit of cytochrome c oxidase (CuCOX), promotes the development of clear cell renal cell carcinoma (ccRCC) through metabolic reprogramming. The authors first observed positive correlation between the level of Cu and disease
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Worrying through the ECM Nat. Cell Biol. (IF 17.3) Pub Date : 2024-12-10 Daryl J. V. David
Cells can migrate through tight spaces by adopting an amoeboid-like motility, characterized by blebbing and minimal proteolytic degradation of the surrounding extracellular matrix (ECM). Driscoll et al. report that amoeboid-like migration can create ECM tunnels by a process they term ‘worrying’, in which large blebs abrade collagen. The authors used melanoma cells, known for amoeboid migration, and
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Regulating chromatin assembly Nat. Cell Biol. (IF 17.3) Pub Date : 2024-12-10 Sabrya Carim
S phase chromatin dynamics safeguard epigenome stability by integrating several processes, including de novo chromatin assembly. Histone chaperone chromatin assembly factor 1, CAF-1, co-ordinates de novo chromatin assembly during DNA replication. Perturbing de novo chromatin assembly in S phase has long-term effects on the epigenome and cell fate; however, the molecular mechanisms are unclear. A study
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Making stronger T cells Nat. Cell Biol. (IF 17.3) Pub Date : 2024-12-10 Stylianos Lefkopoulos
Human induced pluripotent stem cell (iPS cell)-derived chimeric antigen receptor (CAR) T cells hold great therapeutic potential, but often present an immature phenotype compared with αβ T cells used for autologous CAR T cell therapy. A new study shows that H3K9-directed histone methyltransferases G9a–GLP inhibition enables the generation of mature iPS cell-derived T cells. Jing et al. performed a chemical
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Author Correction: Plasma membrane curvature regulates the formation of contacts with the endoplasmic reticulum Nat. Cell Biol. (IF 17.3) Pub Date : 2024-12-02 Yang Yang, Luis A. Valencia, Chih-Hao Lu, Melissa L. Nakamoto, Ching-Ting Tsai, Chun Liu, Huaxiao Yang, Wei Zhang, Zeinab Jahed, Wan-Ru Lee, Francesca Santoro, Jen Liou, Joseph C. Wu, Bianxiao Cui
Correction to: Nature Cell Biology https://doi.org/10.1038/s41556-024-01511-x, published online 17 September 2024.
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Women in Autophagy: an initiative to promote gender parity in science Nat. Cell Biol. (IF 17.3) Pub Date : 2024-12-02 Mericka McCabe, Patricia Boya, Ruey-Hwa Chen, Charleen T. Chu, Maria Isabel Colombo, Laura Delgui, Eeva-Liisa Eskelinen, Maho Hamasaki, Malene Hansen, Congcong He, Marja Jäättelä, Adi Kimchi, Claudine Kraft, Mondira Kundu, Alicia Melendez, Sophie Pattingre, Tassula Proikas-Cezanne, Salwa Sebti, Anna Katharina Simon, Anne Simonsen, Sharon A. Tooze, Maria Ines Vaccaro, Xiaochen Wang, Eileen White, Yan
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The enhancer module of Integrator controls cell identity and early neural fate commitment Nat. Cell Biol. (IF 17.3) Pub Date : 2024-11-26 Yingjie Zhang, Connor M. Hill, Kelsey A. Leach, Luca Grillini, Sandra Deliard, Sarah R. Offley, Martina Gatto, Francis Picone, Avery Zucco, Alessandro Gardini
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FMRP gains mitochondrial fission control Nat. Cell Biol. (IF 17.3) Pub Date : 2024-11-15 Carissa L. Sirois, Soraya O. Sandoval, Xinyu Zhao
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Chromatin remodelling in damaged intestinal crypts orchestrates redundant TGFβ and Hippo signalling to drive regeneration Nat. Cell Biol. (IF 17.3) Pub Date : 2024-11-15 Mardi Fink, Kizito Njah, Shyam J. Patel, David P. Cook, Vanessa Man, Francesco Ruso, Arsheen Rajan, Masahiro Narimatsu, Andreea Obersterescu, Melanie J. Pye, Daniel Trcka, Kin Chan, Arshad Ayyaz, Jeffrey L. Wrana
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FMRP regulates MFF translation to locally direct mitochondrial fission in neurons Nat. Cell Biol. (IF 17.3) Pub Date : 2024-11-15 Adam R. Fenton, Ruchao Peng, Charles Bond, Siewert Hugelier, Melike Lakadamyali, Yi-Wei Chang, Erika L. F. Holzbaur, Thomas A. Jongens
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A systemic effect for liver senescence Nat. Cell Biol. (IF 17.3) Pub Date : 2024-11-13 Kuo Du, David Scott Umbaugh, Rajesh Kumar Dutta, Anna Mae Diehl
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Hepatocellular senescence induces multi-organ senescence and dysfunction via TGFβ Nat. Cell Biol. (IF 17.3) Pub Date : 2024-11-13 Christos Kiourtis, Maria Terradas-Terradas, Lucy M. Gee, Stephanie May, Anastasia Georgakopoulou, Amy L. Collins, Eoin D. O’Sullivan, David P. Baird, Mohsin Hassan, Robin Shaw, Ee Hong Tan, Miryam Müller, Cornelius Engelmann, Fausto Andreola, Ya-Ching Hsieh, Lee H. Reed, Lee A. Borthwick, Colin Nixon, William Clark, Peter S. Hanson, David Sumpton, Gillian Mackay, Toshiyasu Suzuki, Arafath K. Najumudeen