The previous phylogenies of Sordariomycetes by M.E. Barr, O.E. Eriksson and D.L. Hawksworth, and T. Lumbsch and S. Huhndorf, were mainly based on morphology and thus were somewhat subjective. Later outlines by T. Lumbsch and S. Huhndorf, and Maharachchikumbura and co-authors, took into account phylogenetic evidence. However, even these phylogenetic driven arrangements for Sordariomycetes, were somewhat subjective, as the arrangements in trees depended on many variables, such as number of taxa, different gene regions and methods used in the analyses. What is needed is extra evidence to help standardize ranking in the fungi. Estimation of divergence times using molecular clock methods has been proposed for providing additional rational for higher ranking of taxa. Thus, in Sordariomycetes, a divergence period (i.e. 200–300 MYA) can be used as criteria to judge when a group of related taxa evolved and what rank they should be given. In this paper, we provide an updated classification of accepted subclasses, orders of Sordariomycetes and use divergence times to provide additional evidence to stabilize ranking of taxa in the class. We point out and discuss discrepancies where the phylogenetic tree conflicts with the molecular clock.

中文翻译：

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基于系统发育和分子钟证据的Sordariomycetes更新系统发育

ME Barr，OE Eriksson和DL Hawksworth和T.Lumbsch和S.Huhndorf以前对Sordariomycetes的系统发育主要基于形态学，因此有些主观。T. Lumbsch和S. Huhndorf以及Maharachchikumbura及其合着者的后来概述考虑了系统发育证据。但是，即使这些系统进化驱动的拟南芥属真菌的排列也有些主观，因为树木中的排列取决于许多变量，例如分类单元的数量，不同的基因区域和分析中使用的方法。需要额外的证据来帮助标准化真菌的排名。已经提出了使用分子钟法估计发散时间的方法，以便为更高的分类单元排名提供更多的合理性。因此，在Sordariomycetes中，发散期（即 200–300 MYA）可以用作判断一组相关分类单元何时演化以及应给予何种等级的标准。在本文中，我们提供了可接受的子类，Sordariomycetes的顺序的更新分类，并使用发散时间提供了更多证据来稳定分类中的分类单元排名。我们指出并讨论系统发育树与分子时钟冲突的差异。