The earliest fossil record of Phloeocharinae, Angucharcotes thayerae gen. et sp. nov., is reported from mid-Cretaceous Burmese amber. Angucharcotes shares characters with both Phloeocharis-related and Charhyphus-related lineages (e.g., mesoventrite with a median longitudinal carina, temple relatively long, basal tarsomeres unexpanded, pronotal and elytral margins crenulate/serrate, mesocoxae contiguous). The abruptly angulate temples of Angucharcotes are unique in Phloeocharinae. Our discovery of Phloeocharinae from mid-Cretaceous highlights the antiquity of this subfamily.

中文翻译：

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缅甸北部白垩纪中期琥珀中的Phloeocharinae的最早化石记录（鞘翅目：Staphylinidae）

最早的Phloeocharinae化石记录，Angucharcotes thayerae gen。等人。据报道，产自白垩纪中期缅甸琥珀。Angucharcotes与Phloeocharis相关和Charhyphus相关谱系具有共同的特征（例如，具有中位纵向隆突的中腹，太阳穴相对较长，基底节肢节未扩张，前胸和鞘翅边缘具圆齿/锯齿，中壳相连）。Angucharcotes突然成角的寺庙在Phloeocharinae中是独一无二的。我们从白垩纪中期发现的 Phloeocharinae 突出了这个亚科的古代。