The earthworm genus Helodrilus comprises about 20 species with distributions ranging from the Iberian Peninsula to the Caucasus, through Central Europe, the Balkans, Anatolia and the Levant. The species prefer moist habitats and are often found in the mud of river banks, in swamps or in caves. Although a high degree of morphological variability is observed in the taxonomic characters, the genus shares a common feature: the absence of nephridial bladders. In this study, we clarify the phylogeny and taxonomy of the genus Helodrilus using an integrated approach including a multilocus molecular phylogenetic analysis and thorough morphological examination of several species across the putative Helodrilus area of distribution. The findings of this study have led to the genus Helodrilus being split and to the description of two new genera and two new species: Coventina Szederjesi, Sarbu & Csuzdi gen. nov. and Imetescolex movilensis Szederjesi, Sarbu & Csuzdi gen. & sp. nov. The highly homoplasic nature of the absence of nephridial bladders was also revealed.

中文翻译：

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三合一：Helodrilus (Crassiclitellata: Lumbricidae) 属的分子系统发育与两个新属和两个新种的描述

蚯蚓属 Helodrilus 包括大约 20 种，分布范围从伊比利亚半岛到高加索，通过中欧、巴尔干半岛、安纳托利亚和黎凡特。该物种喜欢潮湿的栖息地，经常在河岸的泥浆、沼泽或洞穴中发现。尽管在分类特征中观察到高度的形态变异性，但该属有一个共同特征：没有肾囊。在这项研究中，我们使用一种综合方法阐明了 Helodrilus 属的系统发育和分类学，包括多位点分子系统发育分析和对假定的 Helodrilus 分布区域中的几个物种的彻底形态学检查。这项研究的结果导致了 Helodrilus 属的分裂，并描述了两个新属和两个新物种：Coventina Szederjesi、Sarbu 和 Csuzdi gen。十一月 和 Imetescolex movilensis Szederjesi、Sarbu & Csuzdi gen。＆sp。十一月 还揭示了缺乏肾囊的高度同质性。