Sixty-seven coalified compression specimens of leaves, one attached to a stem, have been found in the St. Mary River Formation (uppermost Campanian–lowermost Maastrichtian, Upper Cretaceous) near Cardston, Alberta, Canada. Leaves have long multi-veined petioles with clasping bases, and an ellipsoid to obovate lamina with a convex to almost flat apex, and a crenate margin with one order of convex-convex chloranthoid teeth with angular sinuses. Tooth apices often end in a small concavity interpreted as a hydathode. Several primary veins enter the lamina in a flabellate manner. The central primary vein branches pinnately while lateral primaries, dichotomize toward the margin producing a reticulodromous pattern of secondaries. Secondaries arising from the lateral primary veins are the result of consistent dichotomies and anastomoses with a concurrent loss of gauge. Tertiary veins are alternate percurrent, and quaternaries form an irregular reticulate fabric. Quaternary veins form polygonal areoles with freely ending veinlets. Some specimens show a polygonal pattern of sub-epidermal aerenchyma like that of many other aquatics. Plants are compared to other fossil aquatic taxa, including those from the St. Mary River Formation and extant members of Ranunculaceae, Menyanthaceae and Nymphaeaceae. The specimens are described as a new amphibious aquatic: Zlatkovia crenulata Rothwell & Stockey gen. et sp. nov., eudicot family incertae sedis. This assemblage meaningfully enriches our understanding of aquatic and adjacent wetland habitats near the close of the Cretaceous, and emphasizes that the structure of modern wetland communities was well established before the end of the Mesozoic.

在加拿大艾伯塔省卡斯顿附近的圣玛丽河组（坎帕阶上部 - 马斯特里赫阶下部，上白垩纪）发现了 67 片煤化压缩叶片标本，其中一个附着在茎上。叶具长多脉叶柄，基部紧握，叶片椭圆形至倒卵形，先端凸至近平，边缘具圆齿，具一阶凸-凸叶绿素齿，具角状鼻窦。齿尖通常以一个小凹面结束，这被解释为一个水囊。几条初级静脉以扇形方式进入椎板。中央初级静脉羽状分支，而侧向初级静脉向边缘二分法，产生网状结构的次级静脉。由侧主静脉产生的继发静脉是一致的二分法和吻合术的结果，同时伴有量规丢失。第三脉为交替的过流，第四脉形成不规则的网状结构。第四纪脉形成具有自由末端细脉的多边形微孔。一些标本显示出与许多其他水生动物一样的表皮下通气组织的多边形图案。将植物与其他化石水生类群进行比较，包括来自圣玛丽河组的那些以及毛茛科、Menyanthaceae 和睡莲科的现存成员。这些标本被描述为一种新的两栖水生动物：一些标本显示出与许多其他水生动物一样的表皮下通气组织的多边形图案。将植物与其他化石水生类群进行比较，包括来自圣玛丽河组的那些以及毛茛科、Menyanthaceae 和睡莲科的现存成员。这些标本被描述为一种新的两栖水生动物：一些标本显示出与许多其他水生动物一样的表皮下通气组织的多边形图案。将植物与其他化石水生类群进行比较，包括来自圣玛丽河组的那些以及毛茛科、Menyanthaceae 和睡莲科的现存成员。这些标本被描述为一种新的两栖水生动物：Zlatkovia crenulata Rothwell & Stockey gen. 等人。11 月，eudicot 家族 incertae sedis。这种组合有意义地丰富了我们对白垩纪末期附近水生和邻近湿地栖息地的理解，并强调了现代湿地群落的结构在中生代末期之前已经很好地建立起来。