Natricine snakes are geographically widespread, species rich (with ~250 extant species) and both morphologically and ecologically diverse. We present a multilocus DNA sequence phylogeny for 249 natricine specimens representing 189 named species, including 69 specimens and 21 species not previously sampled. Our inferred Bayesian and maximum likelihood trees form the basis for evaluations of genus-level classification, historical biogeography, lineage diversification, and dietary, habit and reproductive-mode diversity and evolution, although several, mostly deeper, relationships remain poorly resolved. The optimal trees support natricine origins in Asia, with dispersals to Australo-Melanesia, sub-Saharan Africa (including Seychelles Archipelago, excluding Aldabra), Europe and North Africa and into North and Central America. Viviparity appears to have evolved independently three times in Natricinae but was not significantly associated with an aquatic habit. We found limited associations between habit and diet categories. We propose generic reallocations for four natricine species and highlight other points of uncertainty in natricine classification.

中文翻译：

---

Natricine 蛇的多位点系统发育、自然历史特征和分类 (Serpentes: Natricinae)

Natricine 蛇在地理上分布广泛，物种丰富（约 250 种现存物种）并且在形态和生态上都多样化。我们提出了代表 189 个命名物种的 249 个 natricine 标本的多位点 DNA 序列系统发育，其中包括 69 个标本和 21 个以前未采样的物种。我们推断的贝叶斯树和最大似然树构成了评估属级分类、历史生物地理学、谱系多样化以及饮食、习惯和生殖模式多样性和进化的基础，尽管有几个（大部分是更深层次的）关系仍然很难解决。最佳树种支持 natricine 起源于亚洲，并散布到澳大利亚 - 美拉尼西亚、撒哈拉以南非洲（包括塞舌尔群岛，不包括阿尔达布拉）、欧洲和北非以及北美和中美洲。胎生似乎在 Natricinae 中独立进化了 3 次，但与水生习性没有显着关联。我们发现习惯和饮食类别之间的关联有限。我们建议对四种 natricine 物种进行通用重新分配，并强调 natricine 分类中的其他不确定性。