A new genus of Berothidae (Neuroptera), Speleoberotha gen. nov., with two new species from Brazil, Speleoberotha mineira sp. nov. and Speleoberotha palomae sp. nov., are herein presented. These are the first cave-associated species of Berothidae ever recorded. The new genus shares some characters with the two extant genera of the subfamily Cyrenoberothinae, Cyrenoberotha and Manselliberotha, and other characters are shared with three fossil genera, Microberotha, Protoberotha and Sibelliberotha. This intermediate nature of the new genus was reinforced by the phylogenetic analyses presented here, which recovered Speleoberotha gen. nov. as sister to Sibelliberotha but closely associated with the extant Cyrenoberothinae. In this sense, a new concept of the subfamily Cyrenoberothinae is presented here, now including Speleoberotha gen. nov. and the three fossil genera together with Cyrenoberotha and Manselliberotha. We discuss the biogeographical history of the new concept of Cyrenoberothinae, suggesting that the clade originated in Gondwana. We also provide an identification key for the genera of Cyrenoberothinae.

中文翻译：

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Berothidae（昆虫纲：Neuroptera）的第一个洞穴相关属，以及对 Cyrenoberothinae 亚科的新解释

Berothidae（神经翅目）的一个新属，Speleoberotha gen。十一月，有两个来自巴西的新物种，Speleoberotha mineira sp。十一月 和 Speleoberotha palomae sp。十一月，在此介绍。这是有记录以来第一个与洞穴相关的贝罗特科物种。新属与 Cyrenoberothinae 亚科的两个现存属 Cyrenoberotha 和 Manselliberotha 有一些共同特征，而其他特征与三个化石属 Microberotha、Protoberotha 和 Sibelliberotha 有共同之处。此处介绍的系统发育分析强化了新属的这种中间性质，该分析恢复了 Speleoberotha gen。十一月 作为 Sibelliberotha 的姐妹，但与现存的 Cyrenoberothinae 密切相关。从这个意义上说，这里提出了 Cyrenoberothinae 亚科的新概念，现在包括 Speleoberotha gen。十一月 以及与 Cyrenoberotha 和 Manselliberotha 一起的三个化石属。我们讨论了 Cyrenoberothinae 新概念的生物地理历史，表明进化枝起源于冈瓦纳。我们还提供了 Cyrenoberothinae 属的识别码。