Using a fossilized birth–death model, a new phylogeny of the superfamily Evanioidea (including ensign wasps, nightshade wasps and hatchet wasps) is proposed, with estimates of divergence times for its constitutive families and for corroborating the monophyly of Evanioidea. Additionally, our Bayesian analyses demonstrate the monophyly of †Anomopterellidae, †Othniodellithidae, †Andreneliidae, Aulacidae, Gasteruptiida and Evaniidae, whereas †Praeaulacidae and †Baissidae appear to be paraphyletic. Vectevania vetula and Hyptiogastrites electrinus are transferred to Aulacidae. We estimate the divergence time of Evanioidea to be in the Late Triassic (~203 Mya). Additionally, three new othniodellithid wasps are described and figured from mid-Cretaceous Burmese amber as the new genus Keratodellitha, with three new species: Keratodellitha anubis sp. nov., Keratodellitha basilisci sp. nov. and Keratodellitha kirin sp. nov. We also document a temporal shift in relative species richness between Ichneumonoidea and Evanioidea.

中文翻译：

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包括系统发育中的化石：在尖端测年和化石出生-死亡过程下窥见超科 Evanioidea (Hymenoptera: Apocrita) 的进化

使用化石出生-死亡模型，提出了 Evanioidea 超科（包括少尉黄蜂、茄属黄蜂和斧头黄蜂）的新系统发育，并估计了其组成科的发散时间并证实了 Evanioidea 的单一系。此外，我们的贝叶斯分析表明†Anomopterellidae，†Othniodellithidae，†Andreneliidae，Aulacidae，Gasteruptiida和Evaniidae的单系，而†Praeaulacidae和†Baissidae似乎是并系的。Vectevania vetula 和 Hyptiogastrites electrinus 被转移到 Aulacidae。我们估计 Evanioidea 的发散时间在三叠纪晚期（~203 Mya）。此外，三种新的 thniodellithid 黄蜂在白垩纪中期缅甸琥珀中被描述和推测为新属 Keratodellitha，具有三个新物种：Keratodellitha anubis sp。十一月，Keratodellitha basilisci sp. 十一月 和 Keratodellitha 麒麟 sp。十一月 我们还记录了 Ichneumonoidea 和 Evanioidea 之间相对物种丰富度的时间变化。