Evolutionary models account for either population- or species-level processes but usually not both. We introduce a new model, the FBD-MSC, which makes it possible for the first time to integrate both the genealogical and fossilization phenomena, by means of the multispecies coalescent (MSC) and the fossilized birth–death (FBD) processes. Using this model, we reconstruct the phylogeny representing all extant and many fossil Caninae, recovering both the relative and absolute time of speciation events. We quantify known inaccuracy issues with divergence time estimates using the popular strategy of concatenating molecular alignments and show that the FBD-MSC solves them. Our new integrative method and empirical results advance the paradigm and practice of probabilistic total evidence analyses in evolutionary biology.[Caninae; fossilized birth–death; molecular clock; multispecies coalescent; phylogenetics; species trees.]

中文翻译：

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跨进化时间尺度的分子和形态学的新型综合建模

进化模型解释了种群或物种水平的过程，但通常不能同时考虑两者。我们引入了一种新模型，即 FBD-MSC，它首次可以通过多物种聚结 (MSC) 和化石出生-死亡 (FBD) 过程整合系谱和化石现象。使用这个模型，我们重建了代表所有现存和许多化石犬科动物的系统发育，恢复了物种形成事件的相对和绝对时间。我们使用连接分子比对的流行策略通过发散时间估计来量化已知的不准确问题，并表明 FBD-MSC 解决了这些问题。我们新的综合方法和实证结果推进了进化生物学中概率全证据分析的范式和实践。石化的生死；分子钟；多物种聚结；系统发育学；种树。]