The Cenomanian shallow marine succession in the Southern Galala Plateau of Egypt contains rich benthic fauna and preserves enormous information on how paleo-communities respond to fluctuating environmental conditions. Based on rock and fossil data, the Galala Formation was interpreted to be the result of deposition on a tide-dominated homoclinal ramp. Three main facies associations representing three depositional environments (intertidal, shallow subtidal, and deep subtidal) were recognized. Three distinct macroinvertebrate associations corresponding to these environments and dominated mainly by oysters were quantitatively identified. They show a gradational vertical increase in diversity as well as in the infaunal and mobile species. This trend can be attributed to the Cenomanian sea-level rise, where harsh marginal marine environments in the middle Cenomanian became more stable toward the latest Cenomanian. Sea flooding may have eliminated barriers, enhanced water circulation, and stopped hyperpycnal flow and/or terrigenous flux. Consequently, oxic mesotrophic conditions may have prevailed and siliciclastic starvation may have concentrated fossil materials, and therefore, diversity increased. Based on the vertical and lateral changes of the facies stacking pattern, two main parasequence types were recognized, the first includes deep subtidal deposits changed upwards to shallow subtidal and the second includes shallow subtidal deposits that pass upward into intertidal. Moreover, two and one-half third-order mixed sequences were identified and correlated to their regional and global counterparts.

埃及南部加拉拉高原的 Cenomanian浅海系列包含丰富的底栖动物群，并保存了有关古社区如何应对波动的环境条件的大量信息。根据岩石和化石数据，Galala 组被解释为潮汐主导的同斜斜坡沉积的结果。识别出代表三种沉积环境（潮间带、浅潮下带和深潮下带）的三种主要相组合。三种不同的大型无脊椎动物与这些环境相对应且主要以牡蛎为主的关联被定量地确定。它们显示出多样性以及底栖动物和移动物种的垂直梯度增加。这种趋势可以归因于森诺曼阶海平面上升，森诺曼阶中期恶劣的边缘海洋环境在最近的森诺曼阶变得更加稳定。海洪可能消除了障碍，增强了水循环，并停止超重血流和/或陆源通量。因此，有氧中营养条件可能盛行，硅质碎屑饥饿可能使化石材料集中，因此多样性增加。根据相叠加模式的纵向和横向变化，识别出两种主要的副层序类型，第一种是深潮下沉积向上转变为浅潮下，第二种包括浅潮下沉积向上过渡到潮间带。此外，确定了两个半三阶混合序列并将其与其区域和全球对应序列相关联。