New material from the Upper Devonian (Famennian) Catskill Formation of Pennsylvania, USA, reveals anatomical details previously lacking from the enigmatic fossil species that Newberry (1889, U.S. Geological Survey Monograph 16:1–228) named Holoptychius ? radiatus. The original material (scales and lepidotrichia) was attributed to Holoptychius (Sarcopterygii, Porolepiformes) according to the ridged ornament on the exposed portion of the scales. Scale and cranial anatomy revealed by the new material supports a reassignment of the species to Langlieria (Sarcopterygii, Tristichopteridae). The emended diagnosis of the new combination includes the grooved scale ornament that was noted, though described differently by Newberry, and a posterior oblique pitline groove of the parietal that extends to the caudal margin of the element. Along with Hyneria lindae and Eusthenodon sp. indet., there are now at least three known species of Tristichopteridae from the Catskill Formation. Temporal/spatial separation of these species, along with supporting anatomical characteristics, suggests that tristichopterids commonly filled the ecological niche of top predator in the Famennian-age nonmarine ecosystems of the Catskill Delta Complex. The presence of a Langlieria species in the Catskill Formation lends further support to a biogeographic connection between Pennsylvania, USA, and Belgium during the Famennian Stage of the Upper Devonian.

中文翻译：

---

新材料支持对 Holoptychius 的描述和分类修订？来自美国宾夕法尼亚州上泥盆统卡茨基尔组的 radiatus (Sarcopterygii, Triistichopteridae)

来自美国宾夕法尼亚州上泥盆纪（法门纪）卡茨基尔组的新材料揭示了纽贝里（1889 年，美国地质调查局专着 16:1-228）命名为 Holoptychius 的神秘化石物种以前缺乏的解剖细节。辐射。根据鳞片外露部分的脊状装饰物，原始材料（鳞片和鳞毛）归属于 Holoptychius (Sarcopterygii, Porolepiformes)。新材料揭示的规模和颅骨解剖结构支持将该物种重新分配给 Langlieria（肉翅目，三翅目）。新组合的修正诊断包括注意到的凹槽鳞片装饰，尽管 Newberry 的描述不同，以及延伸到元件尾缘的顶叶后斜坑线凹槽。随着 Hyneria lindae 和 Eusthenodon sp。indet.，现在至少有三种已知的三鳍翅目物种来自卡茨基尔地层。这些物种的时间/空间分离以及支持的解剖特征表明，三翅目通常填补了卡茨基尔三角洲复合体法门时代非海洋生态系统中顶级捕食者的生态位。在上泥盆纪法门阶阶段，卡茨基尔组中存在的 Langlieria 物种进一步支持了宾夕法尼亚州、美国和比利时之间的生物地理联系。表明在卡茨基尔三角洲复合体的法门时代非海洋生态系统中，三叶虫通常占据了顶级捕食者的生态位。在上泥盆纪法门阶阶段，卡茨基尔组中存在的 Langlieria 物种进一步支持了宾夕法尼亚州、美国和比利时之间的生物地理联系。表明在卡茨基尔三角洲复合体的法门时代非海洋生态系统中，三叶虫通常占据了顶级捕食者的生态位。在上泥盆纪法门阶阶段，卡茨基尔组中存在的 Langlieria 物种进一步支持了宾夕法尼亚州、美国和比利时之间的生物地理联系。