ABSTRACT The phylogenetic positions of Cretaceous species of Ceratopogonidae previously placed in the genera Archiculicoides Szadziewski, Protoculicoides Boesel, and Atriculicoides Remm are reappraised in light of synapomorphies. Character states are discussed in detail, supported by new photographs of Protoculicoides depressus Boesel, the description of Protoculicoides revelatus, n. sp., from Burmese amber, and a compilation of previously published illustrations. The recent article by Szadziewski et al. (2016) proposing that Protoculicoides and Atriculicoides are congeneric is shown to be inaccurate. At least three separate lineages are represented by species in these two genera, requiring a new genus, Gerontodacus (type species, G. succineus (Szadziewski)), to include some of them. Archiculicoides, Protoculicoides, Gerontodacus, Adelohelea Borkent and Alautunmyia Borkent remain undetermined to subfamily. As a result of phylogenetic and other taxonomic considerations, the following are new combinations: Gerontodacus krzeminskii (Choufani, Azar, and Nel), Gerontodacus punctus (Borkent), Gerontodacus skalskii (Szadziewski and Arillo), Archiaustroconops andersoni (Szadziewski, Ross, and Giłka), Atriculicoides ciliatus (Borkent), Atriculicoides hispanicus (Szadziewski and Arillo), Atriculicoides sanjusti (Szadziewski and Arillo) and Adelohelea burmitica (Szadziewski and Poinar). The following species are returned to the genera they were assigned to before Szadziewski et al. (2016): Atriculicoides cenomanensis Szadziewski and Schlüter, Atriculicoides dasyheleis Szadziewski, Atriculicoides globosus (Boesel), Atriculicoides incompletus Szadziewski and Schlüter, Atriculicoides macrophthalmus Remm, Atriculicoides sibiricus Szadziewski, Atriculicoides swinhoei (Cockerell), Atriculicoides szadziewskii Pérez-de la Fuente, Delclòs, Peñalver, and Arillo and Atriculicoides taimyricus Szadziewski. A key is provided to all Cretaceous Ceratopogonidae genera.

中文翻译：

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白垩纪咬蠓的系统发育关系，所有已知属的关键（双翅目：蠓科）

摘要 根据突触形态重新评估了先前放置在 Archiculicoides Szadziewski、Protoculicoides Boesel 和 Atriculicoides Remm 属中的 Ceratopogonidae 白垩纪物种的系统发育位置。详细讨论了字符状态，并得到了 Protoculicoides depressus Boesel 的新照片、Protoculicoides revelatus, n. 的描述的支持。sp.，来自缅甸琥珀，以及以前出版的插图的汇编。Szadziewski 等人最近发表的文章。(2016) 提出 Protoculicoides 和 Atriculicoides 是同类的提议被证明是不准确的。这两个属中的物种至少代表了三个独立的谱系，需要一个新属 Gerontodacus（模式种，G. succineus (Szadziewski)）来包括其中的一些。Archiculicoides, Protoculicoides, Gerontodacus, Adelohelea Borkent 和 Alautunmyia Borkent 仍未确定亚科。由于系统发育和其他分类学考虑，以下是新组合：Gerontodacus krzeminskii（Choufani、Azar 和 Nel）、Gerontodacus punctus（Borkent）、Gerontodacus skalskii（Szadziewski 和 Arillo）、Archiaustroconops andersoni、Roszadzid )、Atriculicoides ciliatus (Borkent)、Atriculicoides hispanicus (Szadziewski 和 Arillo)、Atriculicoides sanjusti (Szadziewski 和 Arillo) 和 Adelohelea burmitica (Szadziewski 和 Poinar)。以下物种被归还给 Szadziewski 等人之前分配的属。(2016): Atriculicoides cenomanensis Szadziewski and Schlüter, Atriculicoides dasyheleis Szadziewski, Atriculicoides globosus (Boesel), Atriculicoides incompletus Szadziewski and Schlüter, Atriculicoides macrophthalmus Remm、Atriculicoides sibiricus Szadziewski、Atriculicoides swinhoei (Cockerell)、Atriculicoides szadziewskii Pérez-de la Fuente、Delclòs、Peñalver 和 Arillo 和 Atriculicoides taimyricus Szadziewski。为所有白垩纪 Ceratopogonidae 属提供了一个密钥。