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Balticeler kerneggeri gen. nov., sp. nov., an enigmatic Baltic amber fossil of the ground beetle subfamily Trechinae (Coleoptera, Carabidae)
Deutsche Entomologische Zeitschrift ( IF 1.3 ) Pub Date : 2021-06-02 , DOI: 10.3897/dez.68.66181
Joachim Schmidt , Stephan Scholz , David R. Maddison

Balticeler kerneggerigen. nov., sp. nov., is described based on six fossil specimens preserved in Eocene Baltic amber and imaged using light microscopy and X-ray micro-computed tomography. Based on certain characters observed in the fossil species it is considered a “middle grade” Carabidae, outside of the large family Harpalinae (as it possesses a scrobal seta, the lack of which is a synapomorphy of that subfamily), but possessing four synapomorphies that indicate Balticeler belongs to a large clade of carabids including Harpalinae (anisochaetous Grade B antennal cleaner, conjunct mesocoxae, closed procoxal cavities, and a well-developed external lobe of the metepimeron). This remarkable beetle has several striking features, including lack of externally-visible sexually dimorphic characters, lack of lateral borders on the pronotum, and very long and thin mandibles and maxillae. In combination, these states are unique within Carabidae. We consider the presence of a dorsally completely open aedeagal median lobe as a synapomorphy of the fossil species with the subfamily Trechinae, a pubescent and relatively long second antennomere and a 4+2+2 pattern of umbilicate setae as synapomorphies of the supertribe Trechitae, and a quadrisetose clypeus as a synapomorphy with the Trechitae clade Bembidarenini + Trechini sensu Maddison et al. (2019). As it lacks a synapomorphy of Bembidarenini + Trechini, we propose that it is a member of the stem group of that clade.

中文翻译:

Balticeler kerneggeri gen。十一月,sp。11 月,一种神秘的波罗的海琥珀化石,地甲亚科 Trechinae(鞘翅目,Carabidae)

Balticeler kerneggerigen。十一月,sp。11 月,根据保存在始新世波罗的海琥珀中的六个化石标本进行描述,并使用光学显微镜和 X 射线显微计算机断层扫描成像。根据在化石物种中观察到的某些特征,它被认为是“中级” Carabidae,在大家族 Harpalinae 之外(因为它具有 scrobal 刚毛,缺少的是该亚科的突触),但具有四个突触表明 Balticeler 属于一个大型的 carabids 分支,包括 Harpalinae(anisochaetous B 级触角清洁器,结合 mesocoxae,封闭的 prooxal 腔,以及发育良好的 meepimeron 外叶)。这种非凡的甲虫有几个显着的特征,包括缺乏外部可见的两性异形特征,前胸骨上没有横向边界,以及非常细长的下颌骨和上颌骨。结合起来,这些状态在 Carabidae 中是独一无二的。我们认为背侧完全开放的 aedeagal 中叶作为化石物种与 Trechinae 亚科、短柔毛和相对较长的第二触角和脐状刚毛的 4+2+2 模式作为超级部落 Trechitae 的突触的存在,以及作为与 Trechitae 进化枝 Bembidarenini + Trechini sensu Maddison 等人的突触同形的 quadrisetose clypeus。(2019)。由于它缺乏 Bembidarenini + Trechini 的突触,我们建议它是该进化枝干群的成员。我们认为背侧完全开放的 aedeagal 中叶是化石物种与 Trechinae 亚科、短柔毛和相对较长的第二触角和 4+2+2 脐状刚毛模式的突触,作为超级部落 Trechitae 的突触,以及作为与 Trechitae 进化枝 Bembidarenini + Trechini sensu Maddison 等人的突触同形的 quadrisetose clypeus。(2019)。由于它缺乏 Bembidarenini + Trechini 的突触,我们建议它是该进化枝干群的成员。我们认为背侧完全开放的 aedeagal 中叶作为化石物种与 Trechinae 亚科、短柔毛和相对较长的第二触角和脐状刚毛的 4+2+2 模式作为超级部落 Trechitae 的突触的存在,以及作为与 Trechitae 进化枝 Bembidarenini + Trechini sensu Maddison 等人的突触同形的 quadrisetose clypeus。(2019)。由于它缺乏 Bembidarenini + Trechini 的突触,我们建议它是该进化枝干群的成员。
更新日期:2021-06-02
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