Zethus is the largest genus in Vespidae with over 270 species. It is currently divided into four subgenera: Z. (Zethus), Z. (Zethoides), Z. (Zethusculus) and Z. (Madecazethus). While the last three are restricted to certain biogeographic areas, the first is spread through western and eastern hemispheres. Studies have shown possible phylogenetic incongruence regarding this current division and even raised the possibility of paraphyly in the genus. To evaluate this classification, morphological pasimony analyses under implied weights was carried out, examining external morphology and male genitalia. Analyses showed paraphyly of the genus under various “k” values and paraphyly of two subgenera. Ischnocoelia and Ctenochilus are lowered to subgenera of Zethus. Zethus (Madecazethus) is no longer restricted to Malagasy species and now includes those of the African continent as well. Zethus (Zethastrum)subg. nov. is defined for Oriental representatives. Z. (Zethus) is subdivided into three subgenera: Z. (Zethus), Z. (Didymogastra) and Z. (Wettsteinia). Zethus (Zethusculus) and Z. (Zethoides) are monophyletic and hold their status as subgenera. Three species-groups are Incertae sedis: Z. albopictus, Z. pallidus and Z. spinosus.

中文翻译：

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Zethus Fabricius，1804年（膜翅目，维斯皮科）的星系分析：一种新的亚属分类

Zethus是Vespidae中最大的属，有270多个物种。它当前分为四个子属：Z。（Zethus），Z。（Zethoides），Z。（Zethusculus）和Z.（Madecazethus）。尽管后三个区域仅限于某些生物地理区域，但第一个区域遍布整个西半球和东半球。研究表明，这种当前的分裂可能在系统发育上不一致，甚至增加了属上属的可能性。为了评估该分类，在隐含权重下进行了形态学简化分析，检查了外部形态和男性生殖器。分析显示该属的副系在各种“ k”值下以及两个亚属的副系。坐骨神经鞘病和剑指动物被降低到宙斯的亚属。宙斯（Madecazethus）不再局限于马达加斯加物种，现在也包括非洲大陆的物种。宙斯（Zethastrum）subg。十一月 是为东方代表定义的。Z.（Zethus）分为三个子属：Z.（Zethus），Z。（Didymogastra）和Z.（Wettsteinia）。Zethus（Zethusculus）和Z.（Zethoides）是单系的，并保持其属下的地位。三大类是无翅昆虫：白僵菌Z. pallidus和棘刺Z.。