Newly discovered tetrapod footprints from the middle Permian Pelitic Formation of Gonfaron (Le Luc Basin, Var, France) are described in detail and assigned to the ichnogenus Hyloidichnus. These specimens are very well-preserved, with detailed trackways showing anatomical features, digit drag traces and continuous tail impressions. Together with other Hyloidichnus specimens from the same locality and from the University of Burgundy collections, they allow the identification of the possible trackmakers of Hyloidichnus: small Hyloidichnus footprints are correlated with captorhinomorphs with autopodia similar to Captorhinus whereas large Hyloidichnus footprints might be correlated with larger captorhinomorphs or other ‘basal’ (eventually stem-) moradisaurines. A detailed comparative analysis is proposed to better understand the locomotion style of the trackmakers. Starting from the reconstruction of the stance phase of Captorhinus, we document the swing phase of the fore- and hind-limbs of captorhinomorphs thanks to the continuous digit drag traces associated with Hyloidichnus. A link between gait and track preservation have been highlighted through the analysis of tail impressions and high-resolution 3D models. In all, this study increases our understanding of captorhinomorph locomotion and enhances the integration of both tracks and skeletal remains to highlight the biomechanics of Permian tetrapods.

详细描述了刚法隆中部二叠纪古盆系（法国卢瓦尔盆地）的新发现的四足动物足迹，并将其分配给了鱼鳞Hy。这些标本保存完好，有详细的走线，显示出解剖特征，手指拖曳痕迹和连续的尾巴印痕。加上其他Hyloidichnus来自同一地区和勃艮第藏品大学标本，它们允许的可能trackmakers的标识Hyloidichnus：小Hyloidichnus脚印是相关与captorhinomorphs与autopodia类似大鼻龙属，而大Hyloidichnus脚印可能与较大的captorhinomorphs或其他“基础”（最终是茎）moradisaurine相关。提出了详细的比较分析，以更好地了解田径运动员的运动风格。从站立阶段的重建开始大鼻龙属，我们的文件感谢captorhinomorphs的fore-和后肢的摆动阶段，与之相关联的连续数字拖动的痕迹Hyloidichnus。通过分析尾巴印象和高分辨率3D模型，步态和轨迹保持之间的联系得到了强调。总而言之，这项研究增加了我们对captorhinomorph运动的理解，并增强了轨迹和骨骼遗迹的整合，从而突出了二叠纪四足动物的生物力学。