Recent years have brought undeniable progress in tardigrade taxonomy, and speciose complexes were detected in a number of phylogenetic lineages. The family Echiniscidae is one such lineage; it is one of the most diverse groups of limno-terrestrial tardigrades and can be characterized as having achieved great evolutionary success. In this contribution, using populations representing several species that originated from the Indomalayan region, we reconstructed phylogenetic affinities within Nebularmis, a recently erected genus within the Echiniscus lineage. Nebularmis auratus sp. nov. and Nebularmis burmensis sp. nov. are described from the Eastern Yoma Mountains and the Shan Hills (Myanmar), Nebularmis bhutanensis sp. nov. is described from the Eastern Himalayas (Bhutan), and Nebularmis indicus sp. nov. is described from the foothills of the Western Ghats (Goa, India). Moreover, males are reported in populations of the last two species. All known members of the genus can be phenotypically differentiated based on minute details of their dorsal sculpture and claws. Moreover, a very wide tropical distribution is demonstrated for Nebularmis cirinoi, recorded for the first time from islands of the Malay Archipelago. Furthemore, novel morphological, genetic, and geographic data allowed for the clarification of the generic diagnosis. Currently available data favor a scenario under which Nebularmis evolved in Southeast Asia and later dispersed to other regions of the globe.

中文翻译：

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新的 Indomalayan Nebularmis 物种（Heterotardigrada：Echiniscidae）引发了对其属内多样性的讨论


近年来，缓步动物分类学取得了不可否认的进展，并且在许多系统发育谱系中检测到了物种复合体。 Echiniscidae 科就是这样一个谱系。它是最多样化的湖沼陆生缓步动物类群之一，可以说已经取得了巨大的进化成功。在这篇文章中，我们利用代表来自印度马拉雅地区的几个物种的种群，重建了 Nebularmis 内的系统发育亲缘关系，Nebularmis 是海胆谱系中最近建立的一个属。金星云雾属 sp.十一月和 Nebularmis burmensis sp。十一月Nebularmis bhutanensis sp. 描述于东游马山脉和掸山（缅甸）。十一月描述于东喜马拉雅山（不丹），Nebularmis indicus sp。十一月描述于西高止山脉（印度果阿）山麓。此外，据报道，最后两个物种的种群中有雄性。该属的所有已知成员都可以根据其背部雕塑和爪子的微小细节进行表型区分。此外，Nebularmis cirinoi 的热带分布非常广泛，这是首次在马来群岛岛屿上记录到的。此外，新的形态学、遗传和地理数据可以澄清一般诊断。目前可用的数据支持这样一种情况：Nebularmis 在东南亚进化，后来扩散到全球其他地区。