In recent years, several species that have long been considered to belong in Bacidia s. lat. have been transferred to other genera such as Bellicidia, Bibbya, Scutula, and also to Toniniopsis, accommodating species previously placed in Bacidia and Toninia. One of its widespread species, Toniniopsis subincompta, can be recognized by its thinly granular thallus, dark brown to black apothecia, green epithecium, red-brown hypothecium, and bacilliform ascospores. However, it shows considerable variation in thallus structure, and coloration of apothecia, hypothecium and exciple. We sequenced 20 specimens of T. subincompta to investigate whether there is phylogenetic support for the delimitation of species in accordance with the variability of the observed characters. For phylogenetic analyses, we used newly generated sequence data from the nuclear internal transcribed spacer (nrITS), mitochondrial small subunit (mtSSU) and DNA-directed RNA polymerase II subunit (RPB2). Maximum likelihood and Bayesian inference analyses, as well as three species delimitation programs, provided consistent evidence that T. subincompta forms two separate lineages, to be recognized at the species level. The complex nomenclature of T. subincompta (basionym Lecidea subincompta) shows it to be a synonym of Bellicidia incompta. For the most common taxon previously called Bacidia (Toniniopsis) subincompta, the new combination T. separabilis is made, rather than proposing a conserved type for Lecidea subincompta. Toniniopsis dissimilis is newly described to accommodate the less common taxon. Toniniopsis dissimilis is characterized by a predominantly wrinkled to warted to subsquamulose thallus; generally grey-brown to dark brown apothecia, often with a lighter margin; a dark brown hypothecium, frequently gradually merging into the coloration of the exciple below and the lateral part of the exciple attached to the hymenium; a mostly colourless rim and lateral part of the exciple. The closely related T. separabilis is characterized by a thallus of mostly single or contiguous ±loose granules, often forming short, coralloid, isidium-like bulges; darker apothecia, with a margin mostly of the same colour or darker than the disc; a comparatively thinner hypothecium easily separated from the exciple below. The rim and lateral part of the exciple often contain either a blue, brown or mixed blue-brown colour in the upper part or along the whole margin. Lectotypes of Bacidia vegeta, Lecidea bacillifera f. melanotica and Secoliga atrosanguinea var. affinis (the synonyms of T. separabilis) are selected. Cyanotrophy and the occurrence of albino morphs in T. separabilis are discussed.

中文翻译：

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Toniniopsis subincompta s的形态学和系统发育分析。纬度 (Ramalinaceae, Lecanorales) 在欧亚大陆

近年来，一些长期以来被认为属于杆菌属s。纬度 已转移到其他属，例如贝利西迪亚,比比亚,盾构，并且还连翘属, 容纳先前放置在杆菌属和托尼尼亚. 其分布广泛的物种之一，亚无尾连翘，可以通过其细粒状的菌体、深棕色至黑色的子囊、绿色的上皮、红棕色的子囊和杆状子囊孢子来识别。然而，它显示出菌体结构的相当大的变化，以及 apothecia、hypothecium 和 exciple 的颜色。我们对 20 个样本进行了测序T. subincompta根据观察到的特征的可变性，调查是否有系统发育支持来划分物种。对于系统发育分析，我们使用了来自核内转录间隔区 (nrITS)、线粒体小亚基 (mtSSU) 和 DNA 导向的 RNA 聚合酶 II 亚基的新生成的序列数据。RPB2）。最大似然和贝叶斯推理分析，以及三个物种划界程序，提供了一致的证据表明T. subincompta形成两个独立的谱系，在物种层面被识别。复杂的命名法T. subincompta（基名小鳞甲虫) 表明它是无形贝利西虫. 对于以前称为的最常见的分类单元杆菌属(连翘属)次要的, 新组合T. separabilis是做出来的，而不是提出一个保守的类型小鳞甲虫.异种连翘被新描述以适应不太常见的分类单元。异种连翘特征是主要起皱到疣状到亚鳞状菌体；一般为灰棕色至深棕色的子囊，通常具有较浅的边缘；一个深棕色的小壳，经常逐渐融合到下面的外层和附着在子膜上的外层的外侧部分的颜色中；一个大部分无色的边缘和侧边部分。密切相关的T. separabilis其特征是菌体大多为单个或连续的±松散颗粒，通常形成短的、珊瑚状的、类似isidium的凸起；较深的 apothecia，其边缘的颜色大多与圆盘相同或较暗；一个相对较薄的假体很容易从下面的赋形体中分离出来。外皮的边缘和外侧部分通常在上部或整个边缘包含蓝色、棕色或混合的蓝棕色。选型植物芽孢杆菌,芽孢杆菌F。黑色素和血红球藻变种。亲和（的同义词T. separabilis) 被选中。嗜热症和白化病的发生T. separabilis被讨论。