In this paper, we discuss fitness landscape evolution of permanent replicator systems applying the hypothesis that the specific time of evolutionary adaptation of system parameters is much slower than the time of internal evolutionary dynamics. In other words, we suppose that the extremal principle of Darwinian evolution based on Fisher’s fundamental theorem of natural selection is valid for the steady-states of permanent replicator systems. Various cases illustrating this concept are considered.

中文翻译：

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永久复制系统的适应度优化与进化

在本文中，我们应用系统参数进化适应的特定时间比内部进化动力学的时间慢得多的假设来讨论永久复制系统的适应度景观进化。换句话说，我们假设基于费雪自然选择基本定理的达尔文进化论的极值原理对于永久复制系统的稳态是有效的。考虑了说明这个概念的各种案例。