Sit-and-wait is a common strategy to optimize time and energy devoted to foraging activities. Among Coleoptera, dung beetles are one of the groups that display this behavior. Although diverse aspects of natural history have been reviewed, no agreement exists on which theory may explain this food searching tactic. We performed a systematic literature review about this behavior in dung beetles, analyzing existing studies, and discussing potential explanations. We identified six different hypotheses (resource partitioning, food detection, foraging time, predator’s avoidance, thermoregulation, and resting). Based on the evidence that we found, we support the hypothesis that the main driver of perching behavior is a strategy of spatial segregation. We found reports for several tribes in different biogeographical regions, especially in Neotropical forests. Few studies explore the relationship between perching height and dung beetles’ size. However, recent findings sustained that dung beetles perch at a height proportional to its size, large beetles prefer bigger leaves, and functional groups perch at different heights. Dung beetles observed perching were strictly coprophagous and there is no specific relationship between dial activity and perching behavior. We also found perching records of the three main functional guilds (tunnelers, rollers, and dwellers). Despite several studies of dung beetles perching, there is a lack of experimental analyses. One aspect to analyze in future investigations is the relationship between perching behavior and functional traits. Finally, it is essential to explore and solve some questions that we propose to understand the functional role of this behavior in the structure of assemblages.

中文翻译：

---

粪甲虫（鞘翅目：Scarabaeidae）的栖息行为：稀释高度丰富的组合中的种间竞争的空间隔离机制？

静坐等是优化用于觅食活动的时间和精力的常用策略。在鞘翅目中，甲虫是表现出这种行为的群体之一。尽管已经回顾了自然历史的各个方面，但是对于哪种理论可以解释这种食物搜寻策略尚无共识。我们对about甲的这种行为进行了系统的文献综述，分析了现有研究，并讨论了可能的解释。我们确定了六个不同的假设（资源分配，食物检测，觅食时间，避免捕食，温度调节和休息）。基于我们发现的证据，我们支持以下假设：栖息行为的主要驱动因素是空间隔离策略。我们发现了不同生物地理区域的几个部落的报告，特别是在新热带森林中。很少有研究探索栖息高度与虫大小之间的关系。但是，最近的研究结果表明，be虫的栖息高度与其大小成正比，大甲虫更喜欢较大的叶子，官能团栖息在不同的高度。观察到栖息的粪甲虫是严格共食的，拨号活动与栖息行为之间没有特定的关系。我们还发现了三个主要功能行会（隧道工，压路机和居民）的栖息记录。尽管对several虫栖息有一些研究，但缺乏实验分析。在将来的调查中要分析的一方面是栖息行为与功能特征之间的关系。最后，