We propose and study a three-stage life cycle model of Nemopilema nomurai with a distributed delay to explore the roles of podocyst playing in blooms, and the potential strategies for controlling outbreaks of the giant jellyfish. Our new proposed CPM model is more realistic by considering dormant podocyst, the exclusive form of asexual reproduction of N. nomurai. We investigate the effect of the podocyst production rate and the strobilation rate on the population size, and demonstrate the interannual variation of three phases in the Dalian sea. Our study shows that (1) the long-term dormancy would result in the cardinal number of podocysts multiplying but no discernible effects on a long time dynamics; (2) sensitivity analysis confirms that the recurrence of medusae of N. nomurai is most sensitive to the bottom-up supplement and the key process of the life cycle is strobilation; (3) the effective control strategies are to increase the harvest of ephyrae and to reduce the expansion of substrates.

我们提出和研究的一个三阶段的生命周期模型Nemopilema nomurai与分布式延迟，探讨在华podocyst扮演角色，以及潜在的战略控制巨型水母的暴发。考虑到休眠足囊（野村猪笼草无性繁殖的唯一形式），我们提出的新的CPM模型更加现实。我们调查了荚膜产卵率和增粗率对种群大小的影响，并证明了大连海三个阶段的年际变化。我们的研究表明：（1）长期休眠将导致足囊的基数倍增，但对长时间动态没有明显的影响；（2）敏感性分析证实美杜鹃的复发野村猪笼草对自下而上的补品最敏感，生命周期的关键过程是疏tro。（3）有效的控制策略是增加e的收获并减少底物的膨胀。