Hacquetia epipactis is an understorey species that develops leaves, flowers, and fruits before the canopy layer closes. Its umbels are supported by green bracts. We examined the functional traits of these bracts and basal leaves for specimens, sampled at two altitudes. In this view, their specific leaf/bract area, biochemical, and optical properties were analysed over the course of their development in order to detect differences and changes in both structures. For lowland specimens, specific bract area decreased with age. An increase in UV-absorbing compounds over time was found for both locations. These were negatively related to reflectance in the UV range for basal leaves, whereas for bracts UV reflectance increased with age. We observed an increase in photosynthetic pigment contents during the development of bracts/basal leaves, which resulted in increased absorbance of photosynthetically active radiation for specimens from both altitudes. Detrended correspondence analysis plot based on the reflectance of bracts/basal leaves showed clustering of samples from different age classes with no respect to location. Redundancy analysis revealed high importance of bract/basal leaf biochemistry in explaining reflectance. UV-B- and UV-A-absorbing compounds explained 13% and 8% each, carotenoids 9%, while anthocyanins and chlorophylls explained additional 4% of the reflectance spectra variability. Bracts and basal leaves displayed similar contents of photosynthetic pigments, which indicates their important photosynthetic function for both structures. The development of bracts/basal leaves showed greater influence on the examined parameters in comparison to altitude.

葡萄球菌是一种在冠层关闭之前发育叶片，花朵和果实的下层物种。它的伞形花序由绿色片支撑。我们检查了在两个高度采样的这些片和基生叶的功能特征。以这种观点，在它们的发育过程中分析了它们的比叶/ br片面积，生化和光学特性，以检测两种结构的差异和变化。对于低地标本，特定的片面积随年龄而减少。发现两个位置的紫外线吸收化合物均随时间增加。这些与基叶在紫外线范围内的反射率负相关，而对于for片，紫外线反射率随年龄增加。我们观察到of片/基生叶发育过程中光合色素的含量增加，这导致来自两个高度的标本的光合有效辐射的吸收增加。基于of片/基叶反射率的去趋势对应分析图显示了来自不同年龄类别的样品的聚类，而与位置无关。冗余分析显示，act片/基生叶在解释反射率方面非常重要。吸收UV-B和UV-A的化合物各占13％和8％，类胡萝卜素占9％，而花青素和叶绿素则占反射光谱变异性的4％。ract片和基生叶显示出相似的光合色素含量，表明它们对于两种结构都具有重要的光合功能。与海拔高度相比，of片/基生叶的发育对所检查的参数影响更大。