Abstract Tricalycites Hollick is a distinctive winged propagule that has served as a stratigraphic marker in North American Cretaceous floras. However, its affinities have long been uncertain since the original description of this genus in 1895. Our recent re-examination of specimens from major museum collections and from recent fieldwork has allowed for a revised concept of the morphology and the addition of new details of epidermal anatomy. Twenty three specimens were studied from numerous sites, ranging over a broad area of eastern North America including New Jersey, New York, Massachusetts, Rhode Island, Alabama, and Texas. Tricalycites disseminules have often been called winged fruits, but we cannot eliminate the possibility of them being gymnospermous seeds with a wing derived from an associated envelope or cupule. The type species, T. papyraceus, was previously described by Newberry as having one wing with three lobes or three wings, but we observed only one wing with three prominent lobes. Cuticles of Tricalycites were examined for the first time. The wings lack trichome bases and have paracytic to tetracytic stomata whereas. The new observations facilitate a revised circumscription of the genus Tricalycites but still leave the affinities of this plant open for discussion.

中文翻译：

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Tricalycites 的形态学和表皮解剖学，一种来自北美白垩纪的有翼繁殖体

摘要 Tricalycites Hollick 是一种独特的有翼繁殖体，曾作为北美白垩纪植物群的地层标记。然而，自从 1895 年对该属的最初描述以来，它的亲缘关系一直不确定。我们最近对来自主要博物馆藏品和近期实地考察的标本进行了重新检查，允许修改形态概念并添加表皮的新细节。解剖学。研究了来自多个地点的 23 个标本，范围遍及北美东部的广阔地区，包括新泽西州、纽约州、马萨诸塞州、罗德岛州、阿拉巴马州和德克萨斯州。Tricalycites diseminules 通常被称为有翼果实，但我们不能排除它们是裸子植物种子的可能性，它们的翅膀来自相关的包膜或杯状体。模式种，T。纸莎草，之前被 Newberry 描述为一个翅膀有三个裂片或三个翅膀，但我们只观察到一个翅膀有三个突出的裂片。首次检查了 Tricalycites 的角质层。翅膀没有毛状体基部，有副细胞到四细胞的气孔。新的观察结果促进了对 Tricalycites 属的修订范围，但仍然保留了这种植物的亲和力以供讨论。