ABSTRACT

Heteropachylus Roewer is the type genus of Heteropachylinae Kury, endemic to the Atlantic forest, Brazil. Heteropachyline males bear free tergites at least partially fused to dorsal scutum, besides other sexually dimorphic features common in gonyleptids as a higher ratio of maximum widths of abdominal scute and carapace and stronger armature of legs IV than females. We newly describe Heteropachylus thelydrias sp. nov. from Bahia, Brazil, in which males look like females regarding the shape of dorsal scute and armature of legs IV. Although the presence of polymorphic males is reported to this subfamily with morphs which may have variable sexually dimorphic features, our species appears to lack strongly developed males. Expanding on previous cladistic proposals, we investigated its phylogenetic positioning and found H. thelydrias sp. nov. as sister to a clade containing the remaining species of Heteropachylus (3 MPTs, 316 steps, CI 0.42, RI 0.60). The synapomorphies for Heteropachylus were armature of ocularium, the ectal armature of pedipalp tibia, calcaneus I lighter than astragalus, glans without digitiform dorsal process, and ventral plate with distal border distinctly concave. Although the branch support was low, we refrained from describing a new genus as it would not reflect the phylogenetic information we found.

http://www.zoobank.org/urn:lsid:zoobank.org:pub:B707C01F-B905-4637-A6DF-7597B6C8FC13

摘要

Heteropachylus Roewer 是 Heteropachylinae Kury 的模式属，是巴西大西洋森林特有的。Heteropachyline 雄性具有至少部分融合到背盾板的游离 tergites，除了在 gonyleptids 中常见的其他性二态特征，如腹盾板和甲壳的最大宽度的比例更高，并且腿 IV 的电枢比雌性更强。我们新描述了Heteropachylus thelydriassp。十一月 来自巴西巴伊亚州，其中雄性在背盾的形状和腿 IV 的骨架方面看起来像雌性。尽管据报道该亚科存在多态雄性，其变形可能具有可变的性二态特征，但我们的物种似乎缺乏发育强烈的雄性。扩展先前的分支建议，我们研究了它的系统发育定位并发现了H. thelydrias sp。十一月 作为包含Heteropachylus剩余物种的进化枝的姐妹（3 个 MPT，316 个步骤，CI 0.42，RI 0.60）。Heteropachylus的突触分别为眼膜、胫骨直肠、跟骨I轻于黄芪、龟头无指状背突、腹板远端边缘明显凹入。尽管分支支持率很低，但我们避免描述一个新属，因为它不会反映我们发现的系统发育信息。

http://www.zoobank.org/urn:lsid:zoobank.org:pub:B707C01F-B905-4637-A6DF-7597B6C8FC13