Abstract The sliding and overturning of unconsolidated rubble by hydrodynamic forcing is expected to cause physical damage to settled coral recruits and asexual fragments by scouring and smothering. Yet, few empirical studies have tested the relationship between rubble mobilisation frequency and the survival and growth of these corals. Here, we tested the response of small coral fragments to varying levels of experimental scouring and smothering, proxies for rubble mobilisation impacts, on two coral species with distinct functional traits; Porites rus (robust, encrusting) and Pocillopora verrucosa (sensitive, branching). This study investigated: 1) how coral fragment survival was affected by inversion on rubble or sand substratum over ~2 w (i.e. smothering); and 2) how the survival, growth and tissue properties of fragments were affected by rubble being overturned (i.e. experiencing scouring or abrasion) at different frequencies over ~10 w. Smothering substratum had a profound influence on the probability of fragment survival. For both species, survival was

中文翻译：

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碎石动员对珊瑚碎片存活、部分死亡率和生长的影响

摘要 水动力强迫松散碎石的滑动和倾覆预计会通过冲刷和窒息对定居的珊瑚新兵和无性碎片造成物理损害。然而，很少有实证研究测试碎石移动频率与这些珊瑚的存活和生长之间的关系。在这里，我们测试了小珊瑚碎片对不同程度的实验冲刷和窒息的反应，这是瓦砾动员影响的代理，对两种具有不同功能特征的珊瑚物种；Porites rus（健壮，结壳）和 Pocillopora verrucosa（敏感，分枝）。这项研究调查了：1) 碎石或沙质基质上的倒置如何影响珊瑚碎片的存活率超过约 2 w（即窒息）；2）如何生存，碎片的生长和组织特性受到碎石在约 10 w 的不同频率下翻转（即经历冲刷或磨损）的影响。窒息基质对碎片存活的概率有深远的影响。对于这两个物种，存活率是