Abstract A sampling of angiosperms across a broad spectrum of families and generas was done to determine if angiosperms from the ANA grade basal angiosperms to the Zingiberales of the monocots and Apiales of the eudicots had an endodermis with demonstrable Casparian bands in shoots because the literature has a paucity of demonstrable images, especially for key basal plants. In the Amborellales, Amborella had no stem endodermis, but the aquatic Trithuria of the Nymphaeales had a monostele in which a layer endodermis enclosed its stem axial vascular tissues. Cabomba, Brasenia, and Nuphar of the Nymphaeales had polysteles in which an endodermis of distinct Casparian bands encircled each axial vascular bundle; Nymphaea had either an atactostele or a reduced polystele with only a few axial vascular bundles surrounded by endodermis. Among the magnoliids, Saururus and Houttuynia had monosteles with distinct endodermis Casparian bands. In many monocots, from Acorus, Acorales, to Thalia, Zingiberales, there was a monostele with an endodermis of Casparian bands surrounding a central cylinder of scattered axial vascular bundles. At the base of the eudicots, Ceratophyllum had a vascular core without endodermis. Some members of the Ranunuculales had stem endodermis in a polystelic arrangement. Nelumbo, Proteales, had a monostele of Casparian bands only in stolons. Unusual endodermal and vascular patterns, siphonosteles, were found in Gunnera, Gunnerales. Most eudicots with stem endodermis usually had a monostele or one endodermal layer of Casparian bands only surrounding one ring of vascular bundles, e. g., Myriophyllum, Drosera, Hippuris, Alternanthera, Lysimachia, and Hydrocotyle, but a polystele was present in Justicia. Some leaf petioles and lamina, e. g., Gunnera, Myriophyllum, Hippuris, and Hydrocotyle, had endodermis around their individual vascular bundles. While a stem endodermis was a distinct feature of aquatic or wetland plants, derived from terrestrial ancestors, the presence of stem endodermis by plants at the bases of some major angiosperm lines suggests that it may have been present early in angiosperm evolution.

中文翻译：

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n重新审视被子植物芽中内皮的发生和证据

摘要 对广泛的科和属的被子植物进行了抽样，以确定从 ANA 级基部被子植物到单子叶植物的姜目和双子叶植物的小蜂目中的被子植物是否具有在枝条中具有明显 Casparian 带的内皮层，因为文献中有缺乏可证明的图像，特别是对于关键的基础植物。在 Amborellales 中，Amborella 没有茎内皮，但睡莲目的水生 Trithuria 有一个单柱，其中一层内皮包裹着它的茎轴向维管组织。睡莲目的 Cabomba、Brasenia 和 Nuphar 具有多柱状体，其中不同的里海带的内皮层环绕着每个轴向维管束；Nymphaea 有一个内胚层或一个减少的多柱，只有少数轴向维管束被内皮包围。在木兰科中，Saururus 和 Houttuynia 具有带有明显内皮层 Casparian 带的单星群。在许多单子叶植物中，从 Acorus、Acorales 到 Thalia、Zingiberales，有一个带有 Casparian 带内皮层的单子叶植物，围绕着分散的轴向维管束的中央圆柱体。在真双子叶植物的基部，Ceratophyllum 有一个没有内皮层的血管核心。毛茛目的一些成员具有多列排列的茎内皮。Nelumbo，Proteales，只有在匍匐茎中才有一个 Casparian 乐队的 monostele。在 Gunnera, Gunnerales 发现了不寻常的内胚层和血管模式，即虹吸管。大多数具有茎内胚层的真双子叶植物通常有一个单列或一个内胚层的里氏带，仅围绕一圈维管束，例如，狐尾藻、茅膏菜、马蹄莲、交链孢属、Lysimachia 和 Hydrocotyle，但是在 Justicia 中有一个多石碑。一些叶柄和叶片，例如 Gunnera、Myriophyllum、Hippuris 和 Hydrocotyle，在它们各自的维管束周围具有内皮。虽然茎内皮是源自陆生祖先的水生或湿地植物的一个独特特征，但植物在一些主要被子植物系的基部存在茎内皮表明它可能在被子植物进化的早期就存在。