A serious impediment to the taxonomy of the reticulate whipray Himantura spp. species complex has been the absence of a type specimen for P. Forsskals H. uarnak. Here, reticulate whipray specimens were sampled from the Jeddah region, the assumed type locality of H. uarnak, and characterized genetically at the cytochrome-oxidase subunit 1 (CO1) locus. One of these specimens now in the fish collection of the California Academy of Sciences in San Francisco was designated as neotype. A maximum-likelihood phylogeny of all available CO1 gene sequences from the genus Himantura had the following topology: ((H. leoparda, H. uarnak), (H. undulata, (Himantura sp. 2, (H. australis + Himantura sp. 1))), H. tutul), where H. uarnak haplotypes formed a distinct sub-clade sister to H. leoparda. Based on these CO1 gene sequences, the geographic distribution of H. uarnak includes the eastern Mediterranean, the Red Sea, the East African coast, and the Arabian Sea. Two lineages in the reticulate whipray species complex remain to be named.

中文翻译：

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Forsskål网状鞭子Himantura uarnak的新名称和重新描述

网状鞭子Himantur spp的分类的严重障碍。物种复合体一直缺乏P. Forsskals H. uarnak的类型标本。在这里，网状的鞭状标本是从吉达地区（假定的uarnak型地区）取样的，并在细胞色素氧化酶亚基1（CO1）位点进行了遗传表征。现在，位于旧金山的加利福尼亚科学院鱼类收藏中的这些标本之一被指定为新类型。来自Himantura属的所有可用CO1基因序列的最大似然系统发育具有以下拓扑：（（H. leoparda，H. uarnak），（H. undulata，（Himantura sp。2，（H. australis + Himantura sp。 1））），H。tutul），其中H. uarnak单倍型形成了H. leoparda的一个独特的亚进化姐妹。基于这些CO1基因序列，H的地理分布。uarnak包括地中海东部，红海，东非海岸和阿拉伯海。网状鞭状物种复合体中的两个谱系仍待命名。