Abstract A pteranodontoid pterosaur with anhanguerid affinities (MPSC R 1935) from the Romualdo Formation (Lower Cretaceous, Aptian-Albian), is described here and provides one of the few cases where the ontogenetic stage is established by comparison of skeletal fusion and detailed osteohistological analyses. The specimen comes from the Baixa Grande locality (Araripe Municipality – Ceara State) and is one of the few pterosaurs from the Araripe Basin with precise geographic and stratigraphic data. It consists of a left forelimb, comprising an incomplete humerus, metacarpal IV, pteroid and digits I, II, III, IV, including unguals. This specimen has an estimated maximized wingspan of 7.6 meters, and despite its large dimensions, is considered as an ontogenetically immature individual. Where observable, all bone elements are unfused, such as the extensor tendon process of the first phalanx and the carpal series. The absence of some microstructures such as bone resorption cavities, endosteal lamellae, an external fundamental system (EFS), and growth marks support this interpretation. Potentially, this individual could have reached a gigantic wingspan, contributing to the hypothesis that such large flying reptiles might have been abundant during Aptian-Albian of what is now the northeastern portion of Brazil.

中文翻译：

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巴西东北部阿拉里佩盆地下白垩统 Romualdo 地层大型翼龙的骨组织学和生长模式

摘要 这里描述了来自 Romualdo 组（下白垩纪，Aptian-Albian）的具有 anhanguerid 亲和力的翼齿翼龙 (MPSC R 1935)，并提供了通过比较骨骼融合和详细的骨组织学分析建立个体发育阶段的少数案例之一. 该标本来自 Baixa Grande 地区（阿拉里佩市 - 塞阿拉州），是阿拉里佩盆地为数不多的具有精确地理和地层数据的翼龙之一。它由左前肢组成，包括不完整的肱骨、IV 掌骨、翼骨和指 I、II、III、IV，包括爪。这个标本的最大翼展估计为 7.6 米，尽管它的尺寸很大，但被认为是个体发育不成熟的个体。在可观察到的地方，所有骨骼元素都未融合，如第一指骨和腕骨系列的伸肌腱过程。缺乏一些微结构，如骨吸收腔、骨内膜层、外部基本系统 (EFS) 和生长标记支持这种解释。有可能，这个人可能已经达到了巨大的翼展，这有助于假设这种大型飞行爬行动物在现在巴西东北部的阿普蒂安-阿尔比安期间可能很丰富。