Premise of research. Prominently winged fruits occur in many families of angiosperms today, but they are rare and poorly documented in the mid-Cretaceous. Tricalycites Newberry ex Hollick is a widely distributed winged disseminule from the mid-Cretaceous to Upper Cretaceous sediments of the United States. However, some of the specimens previously attributed to this genus are morphologically distinct, representing a different genus. Methodology. Micro–computed tomography and scanning electron, reflected light, transmitted light, and epifluorescence microscopies were applied for the analyses of compression and impression fossils. Pivotal results. Wireroadia gen. nov. is established on the basis of triwinged fruits from the mid-Cretaceous (Cenomanian) of Alabama. The disseminules include a pedicellate, longitudinally ribbed, globose fruit body and a persistent style. Three prominent longitudinal wings with parallel sides, rounded apexes, and subparallel venation arise from the base of the fruit. The wing epidermis includes prominent trichome bases and paracytic stomatal complexes that are encircled by a ring of radially arranged cells. In addition to the wings, the type species, Wireroadia viccallii sp. nov., shows a pair of prominent antler-like branched spines arising laterally from the base of the fruit. Wireroadia major (Hollick) comb. nov. from the Late Cretaceous (Santonian) of New York, New Jersey, and Massachusetts has larger fruits that lack the branched spines. Both species of Wireroadia are readily distinguished from Tricalycites, which has a miniscule seed body and a single trilobed wing, lacks trichome bases, and bears both paracytic and tetracytic stomata. Conclusions. Wireroadia is interpreted as an extinct genus of eudicot, readily distinguished from Tricalycites. Despite convergent similarities with fruits of Fagales and other extant orders, the more precise systematic position remains uncertain. The presence of both wings and spiny appendages indicates that dispersal might have been by animals, as well as by wind.

中文翻译：

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Wireroadia，来自美国阿拉巴马州和新英格兰白垩纪的带翅果新属

研究的前提。今天，在许多被子植物科中都有明显有翼的果实，但它们在白垩纪中期很少见，而且记录很少。Tricalycites Newberry ex Hollick 是一种广泛分布的有翼散播体，从美国白垩纪中期到上白垩纪沉积物。然而，之前归属于该属的一些标本在形态上是不同的，代表了一个不同的属。方法。显微计算机断层扫描和扫描电子、反射光、透射光和落射荧光显微镜被应用于压缩和印模化石的分析。关键结果。Wireroadia 基因 十一月 是在阿拉巴马州白垩纪中期（森诺曼阶）的三翼果实基础上建立的。散播体包括一个花梗，纵向有棱纹，球状子实体和持久的花柱。三个突出的具有平行侧面的纵向翅、圆形的顶点和近平行的脉络从果实的基部出现。翼表皮包括突出的毛状体基部和被一圈放射状排列的细胞包围的副细胞气孔复合体。除了翅膀，模式种 Wireroadia viccallii sp。11 月，显示了一对突出的鹿角状分枝刺，从果实的基部侧向生长。Wireroadia Major (Hollick) 梳子。十一月 来自纽约、新泽西和马萨诸塞州晚白垩世（桑顿期）的果实较大，但没有分枝刺。Wireroadia 的两个物种很容易与 Tricalycites 区分开来，后者具有微小的种子体和单个三叶翅，缺乏毛状体基部，并带有副细胞和四细胞气孔。结论。Wireroadia 被解释为已灭绝的真双子叶植物属，很容易与 Tricalycites 区分开来。尽管与 Fagales 和其他现存命令的果实有收敛的相似性，但更精确的系统位置仍然不确定。翅膀和多刺附属物的存在表明传播可能是由动物以及风引起的。