Abstract The ichnogenus Kouphichnium and associated ichnofossils attributed to xiphosuran activity are here re-examined from samples collected from the Pennsylvanian-age Steven C. Minkin Fossil Site at the Union Chapel Mine, in Walker County, Alabama, USA. The large sample size offers an unique opportunity to evaluate some Kouphichnium ichnospecies. Thus, the morphological variability resulting from the taphonomic, ethological, taxonomic variability and underprint fallout are evaluated using this large sample set. Three morphotypes have been segregated from the material previously assigned to K. aspodon discovered at this and adjacent sites. Ichnospecies of Kouphichnium identified at the Union Chapel Mine site include: K. lithographicum, K. aspodon and two new ichnospecies (K. atkinsoni and K. minkinensis). Additionally, Kouphichnium-like traces that are associated with “jumper” traces have been previously misinterpreted as Kouphichnium and Selenichnites, respectively, and are excluded from this study, leaving them in open nomenclature as they will be the subject of a subsequent publication. We here redescribe the holotype of K. aspodon and designate lectoparatypes to better define the ichnospecies. New trace makers for some Kouphichnium ichnospecies are hypothesized, in contrast to the traditional xiphosuran attribution.

中文翻译：

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来自美国阿拉巴马州 Steven C. Minkin 古生代足迹遗址（Union Chapel Mines）的 Ichnogenus Kouphichnium 和相关的 Xiphosuran 痕迹：Ichnotaxonomic 和古环境影响

摘要 此处从美国阿拉巴马州沃克县联合教堂矿的宾夕法尼亚时代 Steven C. Minkin 化石遗址收集的样本中重新检查了归因于 xiphosuran 活动的 ichnogenus Kouphichnium 和相关的 ichnofossils。大样本量提供了一个独特的机会来评估一些 Kouphichnium 鱼种。因此，使用这个大样本集评估由栖息地、动物行为学、分类学变异性和底纹脱落引起的形态变异性。三种形态类型已从先前分配给在该地点和邻近地点发现的 K. aspodon 的材料中分离出来。在联合教堂矿场发现的 Kouphichnium 鱼类物种包括：K.lithographicum、K. aspodon 和两个新的鱼类物种（K. atkinsoni 和 K. minkinensis）。此外，与“跳线”痕迹相关的类似 Kouphichnium 的痕迹以前分别被误解为 Kouphichnium 和 Selenichnites，并被排除在本研究之外，使它们处于开放的命名法中，因为它们将成为后续出版物的主题。我们在这里重新描述 K. aspodon 的正模并指定 lectoparatypes 以更好地定义 ichnospecies。与传统的 xiphosuran 归因相反，假设了一些 Kouphichnium 鱼种的新踪迹制造者。aspodon 并指定 lectoparatypes 以更好地定义 ichnospecies。与传统的 xiphosuran 归因相反，假设了一些 Kouphichnium 鱼种的新踪迹制造者。aspodon 并指定 lectoparatypes 以更好地定义 ichnospecies。与传统的 xiphosuran 归因相反，假设了一些 Kouphichnium 鱼种的新踪迹制造者。