During the last few decades, many new ethological categories for trace fossils have been proposed in addition to the original five given by Seilacher. In this article, we review these new groups and present a version of the scheme of fossil animal behaviour originally published by Bromley updated with regard to modern ethological concepts, especially those of Tinbergen. Because some behaviours are more common in certain environments than others, they are useful in palaeoecological reconstructions, forming the original basis of the ichnofacies concept. To simplify, we summarise some ethological categories as previously done by others. However, the tracemaker’s behaviour in some cases is so distinctive that subcategories should be employed, especially in ecological interpretations of certain environments where a special behaviour may be dominant.

中文翻译：

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保存在基质中的动物行为的更新分类

在过去的几十年里，除了 Seilacher 给出的原始五个类别之外，还提出了许多新的痕迹化石行为学类别。在本文中，我们回顾了这些新群体，并展示了最初由 Bromley 发表的关于现代动物行为学概念，尤其是廷伯根的动物行为学概念的更新版本。由于某些行为在某些环境中比其他行为更常见，因此它们在古生态重建中很有用，形成了地貌概念的原始基础。为简化起见，我们总结了其他人以前所做的一些行为学类别。然而，在某些情况下，示踪者的行为是如此独特，以至于应该使用子类别，特别是在某些特殊行为可能占主导地位的特定环境的生态解释中。