Complex interaction among trace fossils are important in understanding organisms niche preference for foraging in ancient rock records. There exist well-recorded examples of compound trace fossils. However, such compound traces rarely exhibit exploitation of burrow wall and fill sediments for feeding. In recent sediments, several examples are well documented for feeding on burrow wall and burrow fills because of nutrient enrichments in them. Such type of complex interactions occur in the shallow marine Pleistocene age sequences exposed in island in Gulf of Kachchh, India. These deposits are known as ‘Shankhodhar Sand Clay Member’ of Dwarka Formation. Trace fossils recorded are Chondrites, Chomatichnus, Gyrolithes, Ophiomorpha, Rhizocorallium, Schaubcylindrichnus, Sinusichnus, Spongeliomorpha, Taenidium and display three Ichnoassemblages. Three types of complex interactions are suggested: (1) Burrow architecture modification, (2) feeding on burrow walls and burrow fills and (3) acute angle branching. Thus, the trace fossil association indicates selective preference of trace maker for their foraging activities in shallow marine Shankhodhar Sand Clay Member of Pleistocene age.

中文翻译：

---

Kachchh 湾更新世 Dwarka 组的地质学分析：示踪行为和返工痕迹

痕量化石之间的复杂相互作用对于理解古代岩石记录中生物体的生态位偏好很重要。存在记录良好的复合痕迹化石的例子。然而，这种复合痕迹很少表现出挖掘洞穴壁和填充沉积物以供取食。在最近的沉积物中，有几个例子很好地记录在洞穴壁和洞穴填充物上，因为它们富含营养。这种复杂的相互作用发生在印度克奇湾岛屿出露的浅海更新世年代序列中。这些矿床被称为 Dwarka 组的“Shankhodhar 砂粘土成员”。记录的痕量化石有球粒陨石、Chomatichnus、Gyrolithes、Ophiomorpha、Rhizocorallium、Schaubcylindrichnus、Sinusichnus、Spongeliomorpha、Taenidium 并显示三个 Ichnosemblages。建议三种类型的复杂相互作用：(1) 洞穴结构修改，(2) 以洞穴壁和洞穴填充物为食，以及 (3) 锐角分支。因此，踪迹化石协会表明踪迹制造者在更新世时代浅海Shankhodhar砂粘土成员中觅食活动的选择性偏好。