Microremains of various sharks, actinopterygians and crocodiles have been recovered from two sites in the Douiret Formation and three sites in the Ain el Guettar Formation in southern Tunisia. The presence of an actinistian is also suggested based on histological study of hemisegments of lepidotrichia. Convergence in dental enameloid microstructure between neoselachian sharks and actinopterygians sharing a tearing dentition is also documented. The vertebrate assemblage of the Douiret Formation suggests a pre-Aptian age for this formation and the presence of Bernissartia in the Ain el Guettar Formation confirms faunal exchange between Africa and Europa during the Early Cretaceous.

中文翻译：

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突尼斯南部早白垩世脊椎动物微遗骸

已从突尼斯南部 Douiret 组的两个地点和 Ain el Guettar 组的三个地点发现了各种鲨鱼、放线鱼类和鳄鱼的微遗骸。根据对鳞状体半节段的组织学研究，也建议存在光化剂。还记录了共享撕裂牙列的新鲨鱼和放线鱼类之间牙釉质微结构的收敛。Douiret 地层的脊椎动物组合表明该地层属于前阿普泰时代，而 Ain el Guettar 地层中 Bernissartia 的存在证实了早白垩世期间非洲和欧罗巴之间的动物群落交换。