Disputing the supposition that ecological competition drives macroevolutionary patterns is now a familiar goal in many fossil biodiversity studies. But it is an elusive goal, hampered by patchy sampling, few assemblage-level comparative analyses, unverified ecological equivalence of clades and a dearth of appropriate statistical tools. We address these concerns with a fortified and vetted compilation of 40190 fossil species occurrences of cyclostome and cheilostome bryozoans, a canonical example of one taxonomically dominant clade being displaced by another. Dramatic increases in Cretaceous cheilostome genus diversification rates begin millions of years before cheilostomes overtake cyclostomes in local species proportions. Moreover, analyses of origination and extinction rates over 150 Myr suggest that inter-clade dynamics are causally linked to each other, but not to changing assemblage-level proportions.

中文翻译：

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当一个化石进化枝取代另一个化石进化枝时，局部优势和进化枝间动态是否存在因果关系？

质疑生态竞争驱动宏观进化模式的假设现在是许多化石生物多样性研究中的一个熟悉的目标。但这是一个难以实现的目标，受到不完整的抽样、很少的组合水平比较分析、未验证的进化枝生态等效性以及缺乏适当的统计工具的阻碍。我们通过对 40190 种圆口和口口苔藓虫化石物种的强化和审查来解决这些问题，这是一个在分类学上占优势的进化枝被另一个进化枝取代的典型例子。白垩纪口口鱼属多样化率的显着增加在口口鱼以当地物种比例超过环口鱼之前数百万年就开始了。而且，