Abstract The clupeomorph genus Armigatus is an extinct group of fish which comprises five species of small marine fishes described from the Late Cretaceous Tethys Sea. Of the five species, A. alticorpus is the least well known and least studied species. The most recent cladistic analyses of the group exclude A. alticorpus due to a lack of sufficient information for the taxon. Here we report new material of Armigatus from the Cenomanian of Hakel, Lebanon. The mass mortality slab preserves multiple clupeomorph fishes including specimens of two species of Armigatus, A. namourensis and A. alticorpus, which have previously been known only from Namoura, Lebanon. The good preservation of skull elements and caudal skeletons of the A. alticorpus specimens provides necessary information for inclusion of this species in a phylogenetic analysis. Results of the phylogenetic reassessment of A. alticorpus indicate that the species had close affinities to the two younger species of the genus, A. oligodentatus and A. dalmaticus. The three species together (A. alticorpus (A. oligodentatus + A. dalmaticus)) belong to a separate lineage from their sister-clade, (A. brevissimus + A. namourensis). Comparative examination of the multiple specimens of each species enabled us to establish a set of morphological traits which allow a more reliable delimitation of the species of Armigatus; this includes number of abdominal scutes, vertebrae, predorsal bones, as well as position of the dorsal fin.

中文翻译：

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黎巴嫩哈克尔上白垩统（森诺曼阶）的 Armigatus (Teleostei, Clupeomorpha, Ellimmichthyiformes) 和 Armigatus alticorpus 新材料的形态变异

摘要 Clupeomorph 属Armigatus 是一种已灭绝的鱼类，包括5 种生活在晚白垩世特提斯海的小型海洋鱼类。在这五个物种中，A. alticorpus 是最不为人所知和研究最少的物种。由于缺乏足够的分类单元信息，该组的最新分支分析排除了 A. alticorpus。在这里，我们报告了来自黎巴嫩哈克尔的塞诺曼尼亚的阿米加图斯的新材料。大量死亡板保存了多种 clupeomorph 鱼，包括两种 Armigatus，A. namourensis 和 A. alticorpus 的标本，它们以前仅在黎巴嫩的纳摩拉已知。A. alticorpus 标本的颅骨元素和尾骨骨骼的良好保存为将该物种纳入系统发育分析提供了必要的信息。A. alticorpus 的系统发育重新评估结果表明该物种与该属的两个年轻物种 A. oligodentatus 和 A. dalmaticus 具有密切的亲缘关系。这三个物种一起 (A. alticorpus (A. oligodentatus + A. dalmaticus)) 属于与其姐妹进化枝 (A. brevissimus + A. namourensis) 不同的谱系。对每个物种的多个标本进行比较检查，使我们能够建立一组形态特征，从而可以更可靠地划定 Armigatus 的物种；这包括腹盾、椎骨、前背骨的数量以及背鳍的位置。alticorpus (A. oligodentatus + A. dalmaticus)) 属于与其姐妹分支 (A. brevissimus + A. namourensis) 不同的谱系。对每个物种的多个标本进行比较检查，使我们能够建立一组形态特征，从而可以更可靠地划定 Armigatus 的物种；这包括腹盾、椎骨、前背骨的数量以及背鳍的位置。alticorpus (A. oligodentatus + A. dalmaticus)) 属于与其姐妹分支 (A. brevissimus + A. namourensis) 不同的谱系。对每个物种的多个标本进行比较检查，使我们能够建立一组形态特征，从而可以更可靠地划定 Armigatus 的物种；这包括腹盾、椎骨、前背骨的数量以及背鳍的位置。