ABSTRACT The Upper Cretaceous (upper Cenomanian–middle Turonian) Akrabou Formation of Asfla, southeast Morocco is renowned for exceptionally preserved, often three-dimensional bony and cartilaginous fish fossils. Teleosts, rare holosteans and chondrichthyans are well known from the so-called ‘Goulmima ichthyological assemblage’, however pycnodonts (Actinopterygii, Pycnodontiformes) have received scant attention and remain undescribed until now. Five nominal species are recognised in the assemblage, including two new forms: Neomesturus asflaensis gen. et sp. nov. and Paranursallia cavini sp. nov. The Italian genus Polazzodus is reported in Morocco for the first time. Specimen taphonomy is examined in the context of substrate consistency and oxygenation of an outer shelf carbonate platform setting. Diversity and ecological disparity of the Goulmima assemblage is revised in regard to trophic partitioning with durophagy seemingly more diverse than previously recognised. Comparisons with other pycnodont-bearing horizons of similar age demonstrate strong faunal affinities between the Asfla pycnodonts and those in the western Tethys and South Atlantic.

中文翻译：

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摩洛哥阿斯弗拉上白垩纪（下图伦阶）Akrabou 组的密齿鱼（Actinopterygii，Pycnodontiformes）

摘要 摩洛哥东南部阿斯弗拉的上白垩纪（上塞诺曼阶 - 中土兰阶）阿克拉布组 (Akrabou Formation) 以保存完好、通常为三维骨软骨鱼化石而闻名。硬骨鱼、稀有全骨类和软骨鱼类在所谓的“古尔米马鱼类学组合”中广为人知，但密齿兽类（Actinopterygii、Pycnodontiformes）却很少受到关注，并且至今仍未被描述。该组合中识别出五个名义物种，包括两种新形式：Neomesturus asflaensis gen。等 十一月 和 Paranursallia cavini sp。十一月 意大利Polazzodus属首次在摩洛哥报道。标本埋藏学是在外陆架碳酸盐平台环境的基质一致性和氧化作用的背景下进行检查的。古尔米玛组合的多样性和生态差异在营养分配方面得到了修正，食硬性动物似乎比以前认识到的更加多样化。与其他类似年龄的密齿兽地层的比较表明，阿斯弗拉密齿兽与特提斯西部和南大西洋的密齿兽之间有很强的动物区系亲缘关系。