Abstract The new ichnospecies of Polykampton georgianum occurs abundantly in a part of the Borjomi Flysch (Upper Paleocene – Lower Eocene) in Georgia, in particular in the 45.3 m thick Ardagani-1 section that is dominated by grey silt- and mudstones having sandstones intercalated. The trace fossil occurs in the middle and upper part of the section. The trace fossil is composed of a median tunnel and side lobes that are filled with material differing from the host sediment. It is interpreted as a sequestrichnion, wherein the tracemaker stored organic-rich mud in the lobes as a food resource for times of food deficiency. The today ferruginization of the median tunnel resulting from the former presence of pyrite suggests strong chemical gradient between its lumen and the surrounding. Chemichnial behaviour is not excluded as nutritional strategy of the worm-like tracemaker (a polychaete?). It occupies a middle tier and is commonly crosscut by Trichichnus, Chondrites and Scolicia. Rare or absent other trace fossils typical of deep-sea fine-grained deposits suggest slight dysoxia. Significant impoverishment of the tiering pattern (only Trichichnus present) and absence of trace fossils in the lower part of the section point to strong dysoxia or episodic anoxia. The trace fossil assemblage is atypical, but in general it resembles the Nereites ichnosubfacies of the Nereites ichnofacies, which characterizes distal parts of the turbiditic depositional systems. The contrast between the dark grey to grey silt- and mudstones deposited during the waning phase of gravitational flows and greenish pelagic mud suggests that the former originated from an organic-rich depositional setting, presumably within the oxygen minimum zone in the upper-slope to outer-shelf domain. Sequestration of the organic-rich mud was a successful adaptation of the Polykampton georgianum producers in this specific environmental setting.

中文翻译：

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格鲁吉亚小高加索古近纪深海复理石矿床中丰富的痕量化石 Polykampton：古生态和古环境影响

摘要 Polykampton georgianum 新物种大量出现在乔治亚州的 Borjomi Flysch（上古新世-下始新世）的一部分，特别是在 45.3 m 厚的 Ardagani-1 剖面中，该剖面以灰色粉砂岩和泥岩夹杂砂岩为主。遗迹化石出现在剖面的中上部。痕迹化石由中间隧道和旁瓣组成，这些旁瓣充满了与宿主沉积物不同的物质。它被解释为隔离，其中示踪剂将富含有机物的泥浆储存在叶瓣中，作为食物缺乏时的食物资源。今天由于黄铁矿的存在而导致的中间隧道的铁锈化表明其管腔和周围环境之间存在强烈的化学梯度。化学行为不排除作为蠕虫状示踪剂（多毛类动物？）的营养策略。它占据中间层，通常被 Trichichnus、球粒陨石和 Scolicia 横切。深海细粒沉积物特有的罕见或不存在的其他痕迹化石表明存在轻微的缺氧症。分层模式的显着贫化（仅存在毛毛虫）和该部分下部没有痕迹化石表明严重的缺氧或间歇性缺氧。微量化石组合是非典型的，但总体上类似于 Nereites ichnofacies 的 Nereites ichnosubfacies，其特征是浊流沉积系统的远端部分。重力流减弱阶段沉积的深灰色至灰色粉砂岩和泥岩与绿色的远洋泥岩之间的对比表明，前者起源于富含有机物的沉积环境，大概在上坡到外坡的最低氧带内。 -货架域。富含有机质泥浆的封存是 Polykampton georgianum 生产商在这种特定环境中的成功适应。