Bathymodiolus Kenk and Wilson, 1985 includes fourteen currently recognized species from deep-sea chemosynthetic environments in the Pacific, Atlantic, and Indian oceans. In this study, phylogenetic analyses of mytilid mussels sampled from seeps along the Costa Rica margin reveal the presence of three new Bathymodiolus species, sampled from depths across ~1000–1900 m. Bathymodiolus billschneideri n. sp., B. earlougheri n. sp., and B. nancyschneiderae n. sp. differ genetically from congeneric species of Bathymodiolus and show some stratification by depth. The depth ranges of Bathymodiolus billschneideri n. sp., B. nancyschneiderae n. sp., and B. earlougheri n. sp. were ~1400–1900 m, 1000–1100 m, and ~1000–1900m, respectively. Bathymodiolus billschneideri n. sp. and B. earlougheri n. sp. were found to be closely related to Bathymodiolus thermophilus Kenk and Wilson, 1985, while B. nancyschneiderae n. sp. exhibited closer relationships to non-East-Pacific taxa. Bathymodiolus thermophilus was the first Bathymodiolus mussel discovered, sampled from a vent along the Galápagos Rift Zone in 1985 and later recorded along much of the East Pacific Rise. This study confirms the presence of B. thermophilus at the Costa Rica margin, representing the first DNA and morphological samples of B. thermophilus at a seep environment. Analysis of habitat evolution suggests that B. thermophilus and its closest relative, B. antarcticus Johnson and Vrijenhoek, 2013, are of seep ancestry.

臭线虫（Bathymodiolus Kenk and Wilson），1985年，包括十四种目前公认的来自太平洋，大西洋和印度洋深海化学合成环境的物种。在这项研究中，从哥斯达黎加边缘的渗流中取样的贻贝贻贝的系统发育分析揭示了存在的三种新的贝氏疟原虫物种，采样深度约为1000-1900 m。斜纹夜蛾。sp。，B. earlougheri n。sp。和B. nancyschneiderae n。sp。从遗传上与水Bath蒲属不同，并且在深度上显示出一些分层。斜纹夜蛾的深度范围。sp。，B. nancyschneiderae n。sp。，和B. earlougheri ñ。sp。分别为〜1400–1900 m，1000–1100 m和〜1000–1900m。斜纹夜蛾。sp。和B.earlougheri n。sp。被发现与嗜热巴斯德氏菌密切相关Kenk和Wilson，1985年，而B. nancyschneiderae n。sp。与非东太平洋分类群的关系更紧密。嗜热巴斯德贻贝是发现的第一只贻贝，自1985年从加拉帕戈斯裂谷带的一个发泄口取样，随后沿东太平洋上升沿大部分地区记录。这项研究证实了哥斯达黎加边缘嗜热芽孢杆菌的存在，代表了嗜热芽孢杆菌的第一个DNA和形态学样品渗水环境中的嗜热芽孢杆菌。对栖息地演变的分析表明，嗜热芽孢杆菌及其最接近的亲缘种南极芽孢杆菌Johnson和Vrijenhoek，2013年具有渗透性。