Abstract Middle Devonian (Givetian) rugose corals from the Holy Cross Mountains (Miloszow), Poland, have been analysed with respect to epibiont colonization patterns. Solitary microconchid tubeworms and colonial bryozoans, hederelloids and ascodictyids numerically dominate on the coral hosts. Moreover, the same taxonomic groups of epibionts dominate both the larger (Heliophyllum, Acanthophyllum, Stringophyllum, Ceratophyllum, Cystiphylloides) and smaller (Thamnophyllum) corallites, indicating the true pattern of epibiosis was achieved. The presence of common bioclaustrated epibionts, constricted and rejuvenated coral calyces, as well as epibionts growing concordantly with the host's growth direction, indicate that at least some corals were colonized during life. The observed low number of overgrowth cases of various epibionts on coral hosts suggests, that competition for hard substrate was negligible, most probably due to abundant coral substrates on muddy sea-bottom of the Givetian sea at Miloszow. This epibiont community is similar to other known Devonian encrusting communities which likely developed in a more nutrient-rich palaeoenvironments and differs markedly from taxonomically-poor assemblages developing in a more oligotrophic conditions. The presence of Rothpletzella encruster, which is considered to be related to green algae, indicates that the Miloszow coral community dwelled within the limits of the photic zone.

中文翻译：

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不同表生生物对皱褶珊瑚的殖民化：波兰圣十字山脉中泥盆纪（Givetian）软底栖息地的优势和同体结壳

摘要 对来自波兰圣十字山脉 (Miloszow) 的中泥盆纪 (Givetian) 皱纹珊瑚进行了表生群落定植模式的分析。单独的微海螺管虫和群居苔藓虫、常石藻和亚丝虫在珊瑚宿主中占主导地位。此外，相同的表生生物分类群在较大的（Heliophyllum、Acanthophyllum、Stringophyllum、Ceratophyllum、Cystiphylloides）和较小的（Thamnophyllum）珊瑚礁中占主导地位，表明实现了表生的真实模式。常见的 bioclaustrated 外生菌、收缩和恢复活力的珊瑚花萼，以及与宿主生长方向一致的外生菌的存在，表明至少有一些珊瑚在生命中被殖民。在珊瑚宿主上观察到的各种表生生物的过度生长情况很少，这表明对硬质基质的竞争可以忽略不计，这很可能是由于米洛佐夫吉维蒂安海泥泞海底上有丰富的珊瑚基质。这个表生群落类似于其他已知的泥盆纪结壳群落，这些群落可能在营养更丰富的古环境中发育，并且与在更贫营养条件下发育的分类学上较差的组合明显不同。被认为与绿藻有关的 Rothpletzella 结壳的存在表明 Miloszow 珊瑚群落居住在光区的范围内。这个表生群落类似于其他已知的泥盆纪结壳群落，这些群落可能在营养更丰富的古环境中发育，并且与在更贫营养条件下发育的分类学上较差的组合明显不同。被认为与绿藻有关的 Rothpletzella 结壳的存在表明 Miloszow 珊瑚群落居住在光区的范围内。这个表生群落类似于其他已知的泥盆纪结壳群落，这些群落可能在营养更丰富的古环境中发育，并且与在更贫营养条件下发育的分类学上较差的组合明显不同。被认为与绿藻有关的 Rothpletzella 结壳的存在表明米洛舒夫珊瑚群落居住在光区的范围内。