We describe the oldest Kalligrammatidae, two distantly related species of different subfamilies from the Lower Jurassic (lower Toarcian) of Germany: Liassopsychops curvatus Bode, 1953, sit. nov. (Liassopsychopinae n. subfam.) and Ophtalmogramma klopschari n. gen. n. sp. (Kallihemerobiinae). They lived in warm and relatively dry conditions. Subfamily Liassopsychopinae includes the Jurassic genera Liassopsychops and Huiyingogramma, sit. nov., which bear forewings characterized by Sc and RA not fused distally; MP, CuA, CuP, and A1 dichotomously branched; and a well-developed central eye-spot. The forewing of Ophtalmogramma n. gen. is most similar to that of Apochrysogramma Yang et al., 2011. The diversity of early Toarcian kalligrammatids indicates a late Triassic–earliest Jurassic origin of the family. A modified classification of the central eye-spots is proposed based on the presence/absence of four main components, i.e., the central pigmented disc (A), swellings (B), ocules (C), and outer rings (D). The function of the swellings is unclear, but they may have served as pheromone containers.

我们描述了最古老的 Kalligrammatidae，这是来自德国下侏罗纪（下 Toarcian）不同亚科的两个远亲物种：Liassopsychops curvatus Bode，1953，坐。十一月 (Liassopsychopinae n. subfam.) 和Ophtalmogramma klopschari n. 将军 n. sp. (Kallihemerobiinae)。他们生活在温暖和相对干燥的环境中。亚科 Liassopsychopinae 包括侏罗纪属Liassopsychops和Huiyingogramma，坐。11 月，其前翅的特征是 Sc 和 RA 未在远端融合；MP、CuA、CuP 和 A1 分枝；和发达的中央眼点。Ophtalmogramma n的前翅。将军 与Apochrysogramma最相似Yang et al., 2011. 早期 Toarcian kalligrammatids 的多样性表明该科起源于晚三叠世 - 最早的侏罗纪。基于四种主要成分的存在/不存在，即中央色素盘 (A)、肿胀 (B)、眼部 (C) 和外环 (D)，提出了中央眼点的改进分类。肿胀的功能尚不清楚，但它们可能充当信息素容器。