A B S T R A C T Multiple-fold hippuritid rudist bivalves –those having infolds of the outer shell layer into the inner shell layer (secondary pillars or rays) other than the three hippuritid pillars– from Eurasia have been referred to genus Pironaea Meneghini. The characters considered in taxonomy caused a somewhat confusing large number of Pironaea species in literature. The study of new material from SE Spain, E Serbia and E and SE Anatolia, stressing the taxonomic relevance of the ontogenetic and intraspecific variability, as well as published material from these or other areas, has resulted in reducing the number of species to three, besides P. milovanovici Kuhn. Thus, a clearer and simpler taxonomic framework of Pironaea species, and their evolutionary relationship, is proposed. P. polystyla (Pirona) is recognized in N and S Italy, Slovenia, Croatia, E and W Serbia, W Bulgaria, Turkey, and UAE, including as junior synonyms P. anatolica Karacabey, P. buseri Pejovic, P. dalmatinica Milovanovic, P. dinarica Milovanovic, P. fruscagorensis Milovanovic, Sladic-Trifunovic, Grubic, P. lisansis Pamouktchiev, P. machnitschi Wiontzek, P. praeslavonica Milovanovic, Sladic-Trifunovic, Grubic, P. sladici Pamouktchiev, and P. slavonica Hilber. P. corrugata (Woodward) is only recognized in E and SE Anatolia and not considered the primitive ancestor of the other species. P. transitoria Milovanovic is recognized in SE Spain, N Italy, E and W Serbia, W Bulgaria, NW Turkey, and Tunisia and includes as junior synonyms P. branislavi Sladic-Trifunovic, P. corrugata nicklesi Philip, P. garlensis Pamouktchiev, P. negrichorensis Pamouktchiev, P. pejovici Pamouktchiev, P. praeyaroslavensis Pamouktchiev, P. timacensis Milovanovic, P. udinensis Pamouktchiev, and P. yaroslavensis Pamouktchiev.

中文翻译：

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Pironaea 种（双壳壳纲：Hippuritidae），除了 P. milovanovici Kühn，来自西班牙东南部、塞尔维亚东部和安纳托利亚东南部，以及来自欧亚大陆上白垩纪的 Pironaea 属的修订版

摘要 来自欧亚大陆的多倍马蹄莲双壳类动物——除三个马蹄莲支柱外，外壳层向内壳层（次生支柱或射线）折叠的双壳贝类——被称为 Pironaea Meneghini 属。分类学中考虑的特征导致文学中大量的 Pironaea 物种有些混乱。对来自西班牙东南部、塞尔维亚东部和安纳托利亚东南部的新材料的研究，强调个体发生和种内变异的分类学相关性，以及来自这些或其他领域的已发表材料，导致物种数量减少到三个，除了 P. milovanovici Kuhn。因此，提出了更清晰、更简单的 Pironaea 物种分类框架及其进化关系。P. polystyla (Pirona) 在意大利北部和南部、斯洛文尼亚、克罗地亚、E 和 W 塞尔维亚、W 保加利亚、土耳其和阿联酋，包括作为初级同义词 P. anatolica Karacabey、P. buseri Pejovic、P. dalmatinica Milovanovic、P. dinarica Milovanovic、P. fruscagorensis Milovanovic、Sladic-Trifunovic、Grubic、 P. lisansis Pamouktchiev、P. machnitschi Wiontzek、P. praeslavonica Milovanovic、Sladic-Trifunovic、Grubic、P. sladici Pamouktchiev 和 P. slavonica Hilber。P. corrugata (Woodward) 仅在安纳托利亚东部和东南部被认可，不被认为是其他物种的原始祖先。P. transtoria Milovanovic 在西班牙东南部、意大利北部、塞尔维亚东部和西部、保加利亚西部、土耳其北部和突尼斯被认可，并包括作为初级同义词 P. branislavi Sladic-Trifunovic、P. corrugata nicklesi Philip、P. garlensis Pamouktchiev、P . negrichorensis Pamouktchiev, P. pejovici Pamouktchiev, P. praeyaroslavensis Pamouktchiev, P.