Abstract The CAMPUS-S model was developed to simulate ground-layer vegetation populations in forest ecosystems and their contribution to carbon and nitrogen dynamics. The model uses several modeling techniques: matrix, l -systems and cellular automata modeling. The genetic species-specific features are defined for each ontogenetic stage (L-systems), while their implementation for certain individuals in the course of modeling depends on external conditions (cellular automata). To implement the variance of the transitions between ontogenetic stages, the matrix modeling method is used. The spatial resolution of the model is 1 cm2 and the time step is one month. The model consists of two submodels: a submodel of the plant populations’ structure, and a submodel of carbon and nitrogen dynamics in plants and soil. The space in the CAMPUS-S model is heterogeneous. The map of the microrelief is dynamically loaded from an external source, according to which the temperature and the moisture of the soil are distributed across space during the simulation. To implement the feedback mechanism between plants and soil, integration with the Romul_Hum model is performed.

中文翻译：

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CAMPUS-S – 森林生态系统中地层植被种群模型及其对碳和氮动态的贡献。一、模型的问题表述与描述

摘要 CAMPUS-S 模型用于模拟森林生态系统中的地层植被种群及其对碳和氮动态的贡献。该模型使用多种建模技术：矩阵、l-系统和元胞自动机建模。每个个体发育阶段（L-systems）都定义了遗传物种特异性特征，而它们在建模过程中对某些个体的实现取决于外部条件（细胞自动机）。为了实现个体发育阶段之间转换的方差，使用矩阵建模方法。模型空间分辨率为1 cm2，时间步长为1个月。该模型由两个子模型组成：一个是植物种群结构的子模型，一个是植物和土壤中碳和氮动态的子模型。CAMPUS-S 模型中的空间是异构的。微地形图是从外部源动态加载的，根据该图，在模拟过程中土壤的温度和水分在空间中分布。为了实现植物和土壤之间的反馈机制，执行了与 Romul_Hum 模型的集成。