The first molecular phylogeny of the tribe Sepidiini is inferred from analyses of DNA sequence data from the following five loci (CAD, wg, COI, COII, 28S rRNA). Bayesian and maximum likelihood analyses were performed on a dataset containing 41 taxa, of which a majority represent Molurina (27) and Phanerotomeina (6). The resulting topologies were used to discuss phylogenetic placement and diagnostic characters of all of the genera representing Molurina. Within the subtribe, the results revealed paraphyly of the genus Psammodes. The P. vialis species-group, currently classified within Psammodes, was recovered as sister to all other Molurina genera. Based on this topology and morphological investigations, a new genus named Toktokkus gen. nov. is established. Within Phanerotomeina, Ocnodes is paraphyletic with regard to Tarsocnodes. In order to restore the monophyly of Ocnodes, the subgenus Chiliarchum stat. nov. is elevated to generic level. Finally, as the homology of female terminalia structures has never been fully assessed for Sepidiini, a comparative study of ovipositor morphology was conducted. As a result, this paper presents the first fully annotated ovipositors for tok-tokkie beetles.

中文翻译：

---

Tok-tokkie甲虫（Tenebrionidae：Molurina，Phanerotomeina）的系统发育的初步见解和Psammodes vialis种组状态的检查

从以下五个位点（CAD，wg，COI，COII，28S rRNA）的DNA序列数据分析推断出Sepidiini部落的第一个分子系统发育。贝叶斯和最大似然分析是在包含41个分类单元的数据集上进行的，其中大多数代表Molurina（27）和Phanerotomeina（6）。所得的拓扑用于讨论代表Molurina的所有属的系统发生位置和诊断特征。在亚部落中，结果揭示了Psammodes属的属植物。该P. vialis种团，目前分类中Psammodes，被恢复为所有其他莫卢里纳属的姐妹。基于此拓扑和形态学研究，一个新属称为Toktokkus属。十一月 成立。在蝶轮虫内部，枕骨相对于Tar骨是共生的。为了恢复八爪鱼的单性，Chiliarchum stat亚属。十一月 提升到通用级别。最后，由于尚未对Sepidiini的雌性末端结构的同源性进行全面评估，因此进行了对产卵器形态的比较研究。结果，本文提出了第一个完全注释的托克-托基甲虫产卵器。