The Plumbaginaceae (non‐core Caryophyllales) is a family well known for species adapted to a wide range of arid and saline habitats. Of its salt‐tolerant species, at least 45 are in the genus Limonium; two in each of Aegialitis, Limoniastrum and Myriolimon, and one each in Psylliostachys, Armeria, Ceratostigma, Goniolimon and Plumbago. All the halophytic members of the family have salt glands and salt glands are also common in the closely related Tamaricaceae and Frankeniaceae. The halophytic species of the three families can secrete a range of ions (Na⁺, K⁺, Ca²⁺, Mg²⁺, Cl⁻, HCO₃⁻, SO₄^2‐) and other elements (As, Cd, Cr, Cu, Fe, Mn, Ni, Pb and Zn). Salt glands are, however, absent in salt‐tolerant members of the sister family Polygonaceae. We describe the structure of the salt glands in the three families and consider whether glands might have arisen as a means to avoid the toxicity of Na⁺ and/or Cl⁻ or to regulate Ca²⁺ concentrations with the leaves. We conclude that the establishment of lineages with salt glands took place after the split between the Polygonaceae and its sister group the Plumbaginaceae.

中文翻译：

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植物中的分泌物结构：榆科植物的起源，进化以及在胁迫耐受性中的作用的教训。

Plumbaginaceae（非核心石竹科）是一个以适应各种干旱和盐碱生境的物种而闻名的科。在其耐盐物种中，至少有45种属于柠檬属。2在每个Aegialitis，Limoniastrum和Myriolimon，各一Psylliostachys，石竹，角柱花属，Goniolimon和白花。该家族的所有盐生植物成员都具有盐腺，并且在紧密相关的Ta科和菊科中也很常见。这三个科的盐生植物可以分泌多种离子（Na ⁺，K ⁺，钙²⁺，镁²⁺，氯^-，HCO ₃ ^-，SO ₄ ^2-）和其它元素（如，镉，铬，铜，铁，锰，镍，铅和锌）。但是，姐妹Poly科的耐盐成员中没有盐腺。我们描述了三个家庭的盐腺的结构，并考虑是否腺体有可能出现的一种手段，以避免娜的毒性⁺和/或Cl ^-或调节钙²⁺浓度的叶子。我们得出的结论是，在gon科及其姊妹科lum科之间分裂后，建立了带有盐腺的血统。