The Ladinian Pseudozilian formation in the central part of the Sava Folds, Slovenia, was deposited in a deeper marine basin situated among the carbonate platforms lining the western margin of the Neotethys. The formation consists of shale and sandstone, hemipelagic limestone, volcanics, and volcanoclastics. At Dole pri Litiji, tuffaceous sandstone contains metre-scale olistoliths dominated by microbialites, solenoporacean algae, sponges, microproblematica and, to a minor extent, corals. Six types of fossil reef associations were recognized based on different proportions of these components. The presence of Ladinella porata Ott, dasycladaceans, and solenoporacean algae suggests the olistoliths originate from within the photic zone, perhaps the platform margin or the upper slope. Similar or even identical associations can be found in other Anisian to early Carnian reefs, suggesting the perseverance of reef associations that established themselves during the Middle Triassic recovery. Rather than replacing the older associations, new associations were added through time. The Anisian–early Carnian reef associations thus probably remained present even after the Late Triassic rise of scleractinian corals to dominance.

中文翻译：

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斯洛文尼亚 Dole pri Litiji 的 Ladinian 碳酸盐岩中的相和化石组合

位于斯洛文尼亚萨瓦褶皱中部的 Ladinian Pseudozilian 地层沉积在一个更深的海盆中，位于新特提斯西缘的碳酸盐台地之间。地层由页岩和砂岩、半远洋灰岩、火山岩和火山碎屑岩组成。在 Dole pri Litiji，凝灰质砂岩包含米级石质岩，主要是微生物岩、龙脑藻、海绵、微问题，以及少量珊瑚。根据这些成分的不同比例，识别出六种类型的化石礁群。Ladinella porata Ott、dasycladaceans 和olenoporacean 藻类的存在表明石质岩起源于光带内，可能是平台边缘或上斜坡。在其他 Anisian 至早期 Carnian 珊瑚礁中可以发现类似甚至相同的关联，这表明在中三叠世恢复期间建立的珊瑚礁组合的持久性。不是替换旧的关联，而是随着时间的推移添加新的关联。因此，即使在晚三叠世石珊瑚上升到主导地位之后，阿尼阶 - 早期卡尼阶珊瑚礁组合可能仍然存在。