Abstract Ediacaran macrofossils are typically preserved in three taphonomic modes: casts/molds in siliciclastic rocks, casts/molds in carbonate rocks, and carbonaceous compressions in black shales. Only a few taxa are known to be preserved in more than one of these taphonomic modes. Flabellophyton is a genus that has been previously reported from lower Ediacaran black shale of the Lantian Formation (635–551 Ma) in South China and upper Ediacaran sandstone of the Ediacara Member (560–550 Ma) in South Australia. Here we report Flabellophyton from upper Ediacaran limestone of the Shibantan Member of the Dengying Formation (551–539 Ma) in South China, making Flabellophyton the only genus that occurs in all three taphonomic modes. We also provide a systematic description of Flabellophyton based on material from the Lantian and Dengying formations in South China, recognizing three morphospecies—F. lantianense, F. typicum sp. nov., and F. obesum sp. nov. Flabellophyton is reconstructed as an erect epibenthic marine organism attached to sandy, carbonate, and muddy substrates. Its phylogenetic affinity remains ambiguous though it was historically interpreted as an algal fossil. The wide environmental and stratigraphic distribution of Flabellophyton allow comparative taphonomic and paleoecological analysis. Taphonomic analysis of Flabellophyton indicates that multiple taphonomic pathways can facilitate the preservation of Ediacaran macrofossils. As a window into Ediacaran paleoecology, Flabellophyton and other Ediacaran fossils played a crucial role in the construction of epibenthic communities in Ediacaran oceans, and helps to understand the ecological migration and evolutionary expansion from deeper to shallower oceans during the Ediacaran Period.

中文翻译：

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三种埋藏模式的故事：保存在石灰岩、黑色页岩和砂岩中的埃迪卡拉纪 Flabellophyton 化石

摘要 埃迪卡拉纪大型化石通常以三种埋藏方式保存：硅质碎屑岩中的铸件/模具、碳酸盐岩中的铸件/模具和黑色页岩中的碳质压缩。已知只有少数分类群以一种以上的这些土埋方式保存。Flabellophyton 是一个先前报道的属，来自华南蓝田组（635-551 Ma）的下埃迪卡拉纪黑色页岩和南澳大利亚埃迪卡拉段（560-550 Ma）的埃迪卡拉纪上层砂岩。在这里，我们报告了来自华南灯影组石板滩段（551-539 Ma）埃迪卡拉纪上层灰岩中的扇藻，使扇藻成为唯一一个同时出现在所有三种埋藏模式中的属。我们还根据来自华南蓝田组和灯影组的材料对扇藻进行了系统描述，识别了三种形态种——F. lantianense, F. typicum sp. 11 月，和 F. obesum sp。十一月 Flabellophyton 被重建为一种直立的底栖海洋生物，附着在沙质、碳酸盐和泥质基质上。尽管它在历史上被解释为一种藻类化石，但它的系统发育亲和力仍然不明确。Flabellophyton 广泛的环境和地层分布允许进行比较的埋藏学和古生态学分析。Flabellophyton 的埋藏分析表明多个埋藏途径可以促进埃迪卡拉纪大型化石的保存。作为了解埃迪卡拉纪古生态学的窗口，