A mid neritic-upper bathyal Ypresian section at Aktulagay, western Kazakhstan, has been analyzed palynologically. A number of key dinoflagellate cyst events are directly calibrated with published calcareous nannofossil data from the same section. The events are used to identify eight dinoflagellate cyst zones from a recently established zonation, used elsewhere in the eastern Peri-Tethys, and to calibrate these zones with the standard nannofossil zonation (NP zones). The events include the lowermost occurrences of Deflandrea oebisfeldensis (∼1%), Dracodinium simile, Eatonicysta ursulae, Dracodinium varielongitudum, Charlesdowniea coleothrypta, Ochetodinium romanum, Charlesdowniea columna, Samlandia chlamydophora, Areosphaeridium diktyoplokum, and Wetzeliella eocaenica. An important regional unconformity separates the Ypresian section from overlying non-calcareous strata with the age-diagnostic species Enneadocysta arcuata, Wetzeliella ovalis, Wilsonidium echinosuturatum, and Rhombodinium draco, indicating the Rhombodinium draco Zone of latest Lutetian–Bartonian age. Based on fluctuations of ecological groups of dinoflagellate cysts, a series of different depositional environments are interpreted and related to the existing sequence stratigraphic model of the section. In most cases dinoflagellate cyst agree with, or supplement, calcareous micro- and nannofossil indications, and support the sequence stratigraphic model. Impagidinium wardii sp. nov. is atypical for the otherwise oceanic genus as it bloomed in a mid-neritic environment. The first cooling at the end of the Early Eocene Climatic Optimum (EECO) is suggested to have caused a strong acme of Eatonicysta ursulae and distinct lowering of the sea level in the NP13 zone. Four new species are formally described: Cribroperidinium cavagnettiae sp. nov., Dracodinium robertknoxii sp. nov., Impagidinium wardii sp. nov., and Samlandia chriskingii sp. nov. The Aktulagay Formation of King et al. (2013) is renamed the Kulsary Formation.

中文翻译：

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来自哈萨克斯坦 Aktulagay 剖面的早中始新世甲藻囊肿

已对哈萨克斯坦西部 Aktulagay 的中浅海-上半深海伊普雷西亚剖面进行孢粉学分析。许多关键的甲藻囊肿事件直接使用来自同一部分的已发布的钙质纳米化石数据进行校准。这些事件用于从最近建立的分区中识别八个甲藻囊肿区，该分区在东特提斯东部的其他地方使用，并使用标准纳米化石分区（NP 区）校准这些区域。这些事件包括最低发生率的 Deflandrea oebisfeldensis (~1%)、Dracodinium simile、Eatonysta ursulae、Dracodinium v​​arielongitudum、Charlesdowniea coleothrypta、Ochetodinium romanum、Charlesdowniea columna、Samlandia chlamydophora、Aerosphaerikizeum。一个重要的区域不整合面将 Ypresian 剖面与上覆的非钙质地层分隔开，这些地层具有年龄诊断物种 Enneadocysta arcuata、Wetzeliella Ovalis、Wilsonidium echinosuturatum 和 Rhombodinium draco，表明最近卢特阶-巴托阶时代的 Rhombodinium draco 带。基于甲藻囊肿生态群的波动，解释了一系列不同的沉积环境，并与该剖面现有的层序地层模型相关联。在大多数情况下，甲藻囊肿符合或补充钙质微化石和纳米化石的适应症，并支持层序地层模型。Impagidinium wardii sp. 十一月 对于其他海洋属来说是不典型的，因为它在中浅海环境中开花。早始新世气候最适（EECO）末期的第一次冷却被认为导致了乌苏拉伊氏藻的强烈顶峰和 NP13 带海平面的明显降低。正式描述了四个新物种： Cribroperidinium cavagnettiae sp。11 月，Dracodinium robertknoxii sp. 十一月，Impagidinium wardii sp。11 月，和 Samlandia chriskingii sp。十一月 King等人的Aktulagay组。(2013) 更名为 Kulsary Formation。