Molecular studies have shown that Liparis s.l. is polyphyletic, with the epiphytic Liparis alliance well separated from the clade representing Liparis in its restricted sense and further forming a phylogenetically heterogeneous assemblage. The generic status of Cestichis, as the key member of the epiphytic Liparis alliance, remains contentious, and the relationships between Cestichis, Stichorkis and Platystyliparis are still unclear and in need of further studies. We herein employ DNA sequence data from nuclear ITS and plastid matK regions to clarify the generic boundaries of Cestichis and its relationship to the allied genera. Results from both Bayesian inference and maximum likelihood analysis congruently indicate that Cestichis as currently circumscribed is not monophyletic. The Blepharoglossum clade, corresponding to commonly recognized sect. Blepharoglossum, forms a monophyletic lineage closely related to the genus Oberonia instead of the core Cestichis clade. A re-evaluation of the diagnostic morphological characters reveals that this lineage represents a morphologically distinct group. Based on these results, Blepharoglossum is removed from Cestichis and elevated to generic status. Our study also sheds light on the homoplastic nature of some diagnostic morphological characters used traditionally in the taxonomy of the Malaxidinae.

中文翻译：

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对 Liparis sl 属（兰花科：Malaxideae）的分子系统发育的新见解，具有新的通用分离物：Blepharoglossum

分子研究表明，Liparis sl 是多系的，附生的 Liparis 联盟与代表 Liparis 的进化枝在其限制意义上完全分离，并进一步形成系统发育异质组合。Cestichis作为附生Liparis联盟的主要成员，其属属地位仍有争议，Cestichis、Stichorkis和Platystyliparis之间的关系尚不清楚，需要进一步研究。我们在此使用来自核 ITS 和质体 matK 区域的 DNA 序列数据来阐明 Cestichis 的通用边界及其与相关属的关系。贝叶斯推断和最大似然分析的结果一致表明，目前限定的 Cestichis 不是单系的。Blepharoglossum 分支，对应于公认的教派。Blepharoglossum 形成与 Oberonia 属密切相关的单系谱系，而不是核心 Cestichis 进化枝。对诊断形态特征的重新评估表明，该谱系代表了一个形态不同的群体。根据这些结果，Blepharoglossum 从 Cestichis 中移除并提升为通用状态。我们的研究还阐明了传统上用于马拉西丁亚科分类学中的一些诊断形态特征的同质性。