Interspecies and intraspecies variation in the use of wooded habitats has been studied for salmonids, but little is known about nonsalmonid species. In this study, we explored spatiotemporal differences in the distribution of three cyprinid species at habitats of varying wood density and at habitats of varying heterogeneity in a Mediterranean intermittent river (Evrotas, southern Greece). Hydroecological data from 843 microhabitats across four sites and in five sampling periods were collected and analysed using generalized linear mixed effects models. Adult chubs, minnowroaches, and minnows and, to a lesser degree, juvenile chubs, preferred complex, wooded habitats, characterized by lower flow velocities, higher water depths, finer substrate, and increased canopy cover, compared with wood‐free and noncomplex habitats. At the advanced stage of the dry period, the deeper, wooded habitats served as drought refugia; all fish groups, except for the adult chub, resorted to these habitats during autumn. In contrast, no such shifts were evident in early summer at the onset of the dry period. In late spring, all fish groups had higher densities at wooded habitats, a shift probably associated with reproduction. Wood density did not affect fish habitat use, as fish densities were similar across wooded habitats. In contrast, habitat heterogeneity significantly affected fish densities; more adult chubs and minnows were observed in complex habitats, whereas juvenile chubs and minnowroaches clearly preferred less complex habitats. We conclude that the targeted maintenance of instream wood structures to ensure habitat heterogeneity is a key‐action for the successful conservation of riverine fish populations.

中文翻译：

---

在流动间歇条件下，木材可用性和栖息地异质性驱动河虾的时空栖息地利用

对于鲑鱼，已经研究了树木繁茂的生境的种间和种内变化，但对非鲑鱼种知之甚少。在这项研究中，我们探讨了在地中海间歇河（Evrotas，希腊南部）的不同木材密度的生境和不同异质性的生境中三种鲤科鱼类物种分布的时空差异。在广义的线性混合效应模型中，收集并分析了来自四个地点和五个采样期的843个微生境的水生态数据。与没有木材和非复杂的生境相比，成年成年的,、 min鱼和min鱼以及较小的幼年成年，更喜欢复杂的树木繁茂的生境，其特征是流速较低，水深较高，底物更细，冠层覆盖率更高。在干旱期的晚期，更深的树木繁茂的栖息地充当了干旱避难所。除成年以外，所有鱼类均在秋季使用这些栖息地。相反，在干燥时期开始的初夏没有明显的这种变化。在春季末期，所有鱼群在树木繁茂的栖息地都具有较高的密度，这一转变可能与繁殖有关。木材密度不影响鱼类栖息地的使用，因为树木茂密的栖息地之间的鱼类密度相似。相反，生境异质性显着影响鱼类密度。在复杂的生境中观察到更多的成年和min鱼，而幼年的chu和min鱼显然更喜欢较不复杂的生境。