Abstract A palynological study of the Upper Cretaceous–lower Paleocene outcrop section at Tattofte (westernmost External Rif, Morocco) has produced a well-preserved palynological assemblage rich in dinoflagellate cysts (dinocysts). For the first time, these assemblages have allowed us to determine the age of this section, to highlight a new Cretaceous–Paleogene boundary, and to make paleobiogeographic interpretations. The samples analyzed yielded 209 dinocyst species and 81 biostratigraphic dinocyst events (first occurrence, or FO, and last occurrence, or LO). We recognized the upper Campanian based on the FOs of Areoligera coronata, Areoligera reticulata, Areoligera senonensis, Cerodinium diebelii and Andalusiella mauthei subsp. mauthei, as well as the LOs of Batiacasphaera solida, Odontochitina costata, Odontochitina porifera, Sentusidinium eisenackii, Trichodinium castanea and Trithyrodinium suspectum. The lower Maastrichtian is determined by the FOs of Isabelidinium bujackii, Palaeocystodinium golzowense and Trithyrodinium evittii, and by the LO of Alterbidinium acutulum, and the upper Maastrichtian is recognized primarily by the FOs of Cerodinium speciosum subsp. speciosum, Disphaerogena carposphaeropsis, Glaphyrocysta perforata and Manumiella seelandica and the LO of Dinogymnium spp. The Danian is recognized on the basis of the FOs of several taxa, including Carpatella cornuta, Danea californica and Senoniasphaera inornata. The succession of the dinocyst events in the uppermost part of the studied interval has revealed a relatively complete record across the Cretaceous–Paleogene boundary. The Cretaceous–Paleogene boundary coincides with a 8 cm-thick reddish layer, which resembles those observed previously in the Cretaceous–Paleogene boundary reference sections in the Mediterranean area, most notably the Cretaceous–Paleogene boundary stratotype at El Kef (Tunisia). The upper Campanian dinocyst assemblage indicates a subtropical to temperate setting, while the Maastrichtian and lower Danian assemblages indicate a subtropical to warm temperate setting. The incursion of a few higher latitude cold-water dinocyst species in these provinces during the latest Campanian and around the Cretaceous–Paleogene boundary may suggest a southward migration of these species, probably related to short cooling episodes and ocean circulation.

中文翻译：

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来自特提斯西南边缘的坎帕尼亚至大年甲藻囊肿组合（Tattofte 剖面，西外里夫，摩洛哥）：生物地层学和古生物地理解释

摘要 对塔托夫特（摩洛哥最西端的外部裂谷）上白垩统-下古新世露头部分的孢粉学研究产生了一个保存完好的富含甲藻囊肿（甲藻）的孢粉学组合。这些组合首​​次使我们能够确定该剖面的年龄，突出一个新的白垩纪-古近纪边界，并进行古生物地理解释。分析的样本产生了 209 种双囊壳物种和 81 个生物地层双囊壳事件（首次出现，或 FO，最后一次出现，或 LO）。我们根据 Areoligera coronata、Areoligera reticulata、Areoligera senonensis、Cerodinium diebelii 和 Andalusiella mauthei subsp。mauthei，以及 Batiacasphaera solida、Odontochitina costata、Odontochitina porifera 的 LO，Sentusidinium eisenackii、Trichodinium castanea 和 Trithyrodinium guessum。下马斯特里赫特由 Isabelidinium bujackii、Palaeocystodinium golzowense 和 Trithyrodinium evittii 的 FO 以及 Alterbidinium acutulum 的 LO 决定，而上马斯特里赫特主要由 Cerodinium speciosum subsp. 的 FO 识别。speciosum、Disphaerogena carposphaeropsis、Glaphyrocysta perforata 和 Manumiella seelandica 以及 Dinogymnium spp 的 LO。Danian 是根据几种分类群的 FO 来识别的，包括 Carpatella cornuta、Danea californica 和 Senoniasphaera inornata。研究区间最上部的双胞胎事件的连续性揭示了跨越白垩纪-古近纪边界的相对完整的记录。白垩纪-古近纪边界与 8 厘米厚的红色层重合，这类似于先前在地中海地区的白垩纪-古近纪边界参考剖面中观察到的那些，最显着的是 El Kef（突尼斯）的白垩纪-古近纪边界层型。上坎帕尼亚双胞囊组合表明亚热带至温带环境，而马斯特里赫特和下大尼阶组合表明亚热带至暖温带环境。在最近的坎帕阶和白垩纪-古近纪边界附近，这些省份的一些高纬度冷水双胞藻物种的入侵可能表明这些物种向南迁移，可能与短暂的冷却事件和海洋环流有关。最显着的是 El Kef（突尼斯）的白垩纪-古近纪边界层型。上坎帕尼亚双胞囊组合表明亚热带至温带环境，而马斯特里赫特和下大尼阶组合表明亚热带至暖温带环境。在最近的坎帕阶和白垩纪-古近纪边界附近，这些省份的一些高纬度冷水双胞藻物种的入侵可能表明这些物种向南迁移，可能与短暂的冷却事件和海洋环流有关。最显着的是 El Kef（突尼斯）的白垩纪-古近纪边界层型。上坎帕尼亚双胞囊组合表明亚热带至温带环境，而马斯特里赫特和下大尼阶组合表明亚热带至暖温带环境。在最近的坎帕阶和白垩纪-古近纪边界附近，这些省份的一些高纬度冷水双胞藻物种的入侵可能表明这些物种向南迁移，可能与短暂的冷却事件和海洋环流有关。