Abstract Scombroclupea javieri sp. nov. is erected based on the description of well-preserved specimens collected in the lower Cenomanian marine outcrop at the El Chango quarry, near Ocozocoautla de Espinosa, Chiapas State, southeastern Mexico. This new species shows the distinctive feature of the superorder Clupeomorpha, the presence of the abdominal scute series, as well as the diagnostic characters of Scombroclupea, as the lack of the dorsal scutes and the presence of two supramaxillae, the comb-like dentition in the maxilla, the horizontal laminae in the anterior caudal vertebrae, the flat epineurals and epipleural in caudal region of the trunk, and the finlets present along the anal fin, and differs from the other three nominal species of this genus in meristic data of the fins and vertebrae. Scombroclupea was previously thought as an incertae sedis members of the order Clupeiformes; however, S. javieri displays two primitive features never recorded before in this clupeomorph order and commonly observed in the order Ellimmichthyiformes, this new species has a large beryciform foramen piercing the anterior ceratohyal and its parietal bones are sutured to each other. The review of specimens of S. occidentalis and S. macrophthalma confirms that these species share the presence of a large beryciform foramen in the anterior ceratohyal. These characters make it difficult to include Scombroclupea as a member of any of the orders Clupeiformes and Ellimmichthyiformes. This record of Scombroclupea, the oldest in America and the second in Mexico, widens the geographical Cenomanian distribution of the genus along the Tethys Sea, from Middle East to southern North America.

中文翻译：

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Scombroclupea javieri sp. 11 月，一种神秘的 Cenomanian clupeomorph 鱼（Teleostei，Clupeomorpha）来自墨西哥东南部恰帕斯州 Ocozocoautla Cintalapa 组的海洋沉积物

摘要 Scombroclupea javieri sp。十一月 是根据对在墨西哥东南部恰帕斯州 Ocozocoautla de Espinosa 附近的 El Chango 采石场的 Cenomanian 海底露头收集的保存完好的标本的描述而建造的。这个新种显示了超级目 Clupeomorpha 的显着特征，存在腹甲系列，以及 Scombroclupea 的诊断特征，因为没有背甲和存在两个上颌骨，在上颌骨、前尾椎骨的水平椎板、躯干尾部区域的扁平神经外膜和胸膜外膜，以及沿着臀鳍存在的小鳍，在鳍的分生数据上与该属的其他三个名义种不同椎骨。Scombroclupea 以前被认为是 Clupeiformes 的一个不确定的 sedis 成员。然而，S. javieri 显示了两个以前从未记录在此clupeomorph 目中的原始特征，并且在Ellimmichthyiformes 目中常见，这个新物种有一个巨大的beryciform 孔，穿透前角板，顶骨相互缝合。对 S. occidentalis 和 S. macrophthalma 标本的审查证实，这些物种在前角玻璃中都有一个大的铍状孔。这些特征使得很难将 Scombroclupea 作为 Clupeiformes 和 Ellimmichthyiformes 目中的任何一个的成员。Scombroclupea 是美洲最古老、墨西哥第二的记录，它扩大了该属沿特提斯海的地理分布，